
Belongs within: Nassellaria.
The Acanthodesmiidae are a group of radiolarians known from the Eocene to the present (Afanasieva et al. 2005).
Characters (from Afanasieva et al. 2005): Cephalis or its derivatives present. Differentiation in direction of main heteropolar axis indistinct even in inner skeletons. Spines L developed more weakly than spine D and l. External apophyses, compared with feet formed by spines L absent. Ante- and postcephalic lobes of cephalis developed very strongly. May be developed mitral–galear structures.
<==Acanthodesmiidae [Acanthodesmiacea, Acanthodesmida, Paratympanidae, Trissocyclidae, Tympanida]AA05 | i. s.: Tristephanium Haeckel 1881 (n. o.)H04, AA05 | `--T. dimensivumH04 | Octotympanum Haeckel 1887 (n. o.)H04, AA05 | `--O. cervicorneH04 | Circotympanum Haeckel 1887H04, AA05 | `--C. octogoniumH04 | Lithocubus Haeckel 1881H04, AA05 | `--L. astragalusH04 | Paratympanum Haeckel 1881AA05 |--Paradictyum Haeckel 1881 [Paradictyinae]AA05 | `--P. paradoxumAA05 |--PerispyridinaeAA05 | |--Perispyris Haeckel 1881AA05 | |--Tricolospyris Haeckel 1881AA05 | |--Lithotympanum Haeckel 1881AA05 | | `--L. tuberosumH04 | `--Microcubus Haeckel 1881AA05 | `--M. zonariusH04 |--Circospyridinae [Protympaniinae]AA05 | |--Amphispyris Haeckel 1881AA05 | |--Archicircus Haeckel 1887AA05 | |--Circospyris Haeckel 1881AA05 | |--Sphaerospyris Haeckel 1887AA05 | |--Spyridobotrys Haeckel 1862AA05 | |--Toxarium Haeckel 1887AA05 | |--Protympanium Haeckel 1881AA05 | |--Lithocircus Müller 1856AA05 | | `--L. magnificusH04 | `--Toxidiella Loeblich & Tappan 1961LT61 [=Toxidium Haeckel 1887 non LeConte 1860AA05] | `--*T. cordatum (Haeckel 1887) [=*Toxidium cordatum, Toxarium (*Toxidium) cordatum]LT61 `--AcanthodesmiinaeAA05 |--Acrocubus Haeckel 1881AA05 |--Dipocoronis Haeckel 1881AA05 |--Eucoronis Haeckel 1881AA05 |--Tetracubus Haeckel 1887AA05 |--Tricyclidium Haeckel 1881AA05 |--Tripocubus Haeckel 1887AA05 |--Trissocyrcus Haeckel 1881AA05 |--Tympanomma Haeckel 1887AA05 |--Zygostephanus Haeckel 1862AA05 |--Semantis Haeckel 1887AA05 | `--S. sigillumH04 |--Trissocyclus Haeckel 1881 [incl. Trissocircus Haeckel 1881]AA05 | `--T. sphaeridiumH04 |--Tympanidium Haeckel 1881AA05 | `--T. foliosumH04 |--Acanthodesmia Müller 1856AA05 | |--A. coronaH04 | `--A. vinculataAA05 `--Tympaniscus Haeckel 1887AA05 |--T. quadrupesH04 `--T. tripodiscusH04
*Type species of generic name indicated
References
[AA05] Afanasieva, M. S., E. O. Amon, Y. V. Agarkov & D. S. Boltovskoy. 2005. Radiolarians in the geological record. Paleontological Journal 39 (Suppl. 3): S135–S392.
[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.
[LT61] Loeblich, A. R., Jr & H. Tappan. 1961. Remarks on the systematics of the Sarkodina (Protozoa), renamed homonyms and new and validated genera. Proceedings of the Biological Society of Washington 74: 213–234.