Achatinina

Opeas pyrgula, copyright H. Zell.

Belongs within: Heterobranchia.
Contains: Streptaxidae, Achatininae, Stenogyrinae, Subulininae.

The Achatinina are a clade of snails united by molecular phylogenetic analysis, members of which have genitalia with a greatly reduced or absent stimulator. The carnivorous Streptaxoidea, characterised by a radula with lanceolate teeth, form the sister group of the remaining species in the Achatinoidea.

<==AchatininaBR17
|--StreptaxoideaBR17
| |--StreptaxidaeBR05
| `--Diaphera Albers 1850 [Diapheridae]BR17
| `--*D. cumingiana (Pfeiffer 1845) [=Cylindrella cumingiana]BR17
`--Achatinoidea [Achatinacea]BR05
|--Micractaeon Verdcourt 1993 [Micractaeonidae]BR05
| `--*M. kakamegaensis Verdcourt 1993BR17
|--Aillya Odhner 1927BR05 [Aillyida, AillyidaeBR17, Aillyoidea]
| `--*A. camerunensis Odhner 1927BR17
|--Ferussaciidae [Ferrusacidae, Ferussaciinae]SS10
| |--CoilosteleTTE93
| |--Ferussacia Risso 1826 [=Ferrussacia (l. c.)]BR05
| | |--*F. gronoviana Risso 1826BR17
| | |--F. folliculum (Schröter 1784)NB19
| | `--F. subcylindricaW79
| `--Geostilbia Crosse 1867SS10
| `--G. gundlachi (Pfeiffer 1850) (see below for synonymy)SS10
`--Achatinidae [Achatinae, Achatinida, Subulinidae, Subulinoidea]SS10
| i. s.: Leptocala mollicellaFGN07
| OreohomorusP02
| SubulonaFGN07
| |--S. bumbaensis (Spence 1923) [=Homorus (Subulona) bumbaensis]PB27
| `--S. decollata (Morelet 1873)FGN07
| StriosubulinaFGN07
| DictyoglessulaFGN07
| OleataFGN07
| PileataFGN07
| Eremopeas Pilsbry 1906SS10
| |--*E. interioris (Tate 1894)SS10 [=Stenogyra interiorisSS10, Pseudopeas (Eremopeas) interiorisS88]
| `--E. tuckeri (Pfeiffer 1846)SS10 (see below for synonymy)
| Prosopeas achatinaceum (Pfeiffer 1846) (see below for synonymy)S88
| Allopeas Baker 1935SS10
| |--*A. gracile (Hutton 1834)SS10 (see below for synonymy)
| |--A. clavulinus (Potiez & Michaud 1838)SS10
| `--A. micrus (d’Orbigny 1835) [=Lamellaxis (Allopeas) micrus]J68
| Paropeas Pilsbry 1906SS10
| |--*P. acutissimum (Mousson 1857) [=Bulimus acutissimus]SS10
| `--P. achatinaceum (Pfeiffer 1846)SS10
| EutomopeasBR17
| LeptinariaBR17
| |--L. elisae Tristram 1861C64
| |--L. emmelinae Tristram 1861C64
| `--L. lamellata [=Tornatellina (Leptinaria) lamellata]A66
| ArchachatinaPB27
| ColumnaPB27
| |--C. (Rhodea) californica [=Achatina californica]C64
| `--C. flammeaD66
| PseudachatinaPB27
| AtopocochlisPB27
| Pseudotrochus jansseni Dupuis & Putzeys 1922PB27
| ThomeaPB27
| KrapfiellaPB27
| MabiliellaPB27
| MetachatinaPB27
| CochlitomaPB27
| HypolysiaPB27
| XerocerastusPB27
| ZootecusPB27
| ChilonopsisPB27
| Neosubulina (Ischnocion) triptyxT68
|--AchatininaeBR05
|--StenogyrinaeBR05
|--SubulininaeBR17
|--Trichodina Ancey 1888BR17 [=Petriola Dall 1905BR17; PetriolinaeBR05]
| `--*T. marmorea (Reeve 1850) [=Achatina marmorea, *Petriola marmorea]BR17
|--Rumina Risso 1826 [Ruminidae, Rumininae]BR05
| `--*R. decollata (Linnaeus 1758)BR17 [=Helix decollataBR17, Bulimus decollatusG40]
|--Coeliaxis Adams & Angas 1865 [=Caeliaxis (l. c.); Caeliaxidae, Coeliaxinae]BR05
| `--*C. layardi (Adams & Angas 1865) [=Subulina layardi]BR17
|--Pyrgina Greef 1882BR05 [PyrgininaeBR17]
| `--*P. umbilicata Greef 1882BR17
|--Cryptelasmus Pilsbry 1907BR05 [CryptelasminaeBR17]
| `--*C. canteroianus (Pfeiffer 1857) [=Balea canteroiana]BR17
|--Thyrophorella Greef 1882BR05 [Thyrophorellidae, ThyrophorellinaeBR17, Thyrophorelloidea]
| `--*T. thomensis Greef 1882BR17
|--Cecilioides Férussac 1814BR05 (see below for synonymy)
| |--*C. acicula (Müller 1774)BR17 (see below for synonymy)
| |--C. eulimaCC06
| |--C. gundlachi [incl. Caecilioides spencei Dupuis & Putzeys 1922]PB27
| |--C. petitiana (Benoit 1862)NB19
| `--C. stuhlmanniPB27
|--Opeas Albers 1850 [Opeatinae]BR05
| |--*O. goodallii (Miller 1822)BR17 (see below for synonymy)
| |--O. adamsiT68
| |--O. minutumE99
| |--O. pyrgula Schmacker & Boettger 1891S88
| |--O. thomasi Connolly 1923PB27
| `--O. venustumPB27
`--Glessulinae [Glessulidae]BR17
|--TortaxisBR17
|--Glessula Martens 1860BR05
| `--*G. gemma (Reeve 1850) [=Achatina gemma]BR17
`--Rishetia Godwin-Austen 1920BR17, BR05 [Rishetiinae]
`--*R. tenuispira (Benson 1836) [=Achatina tenuispira]BR17

*Allopeas gracile (Hutton 1834)SS10 [=Bulimus gracilisSS10, Lamellaxis (Allopeas) gracilisS88, Opeas gracileS88, Stenogyra gracilisTC89; incl. O. ardelio Iredale 1941SS10, O. oparanum (Pfeiffer 1846)S88]

Cecilioides Férussac 1814BR05 [=Caecilianella Bourguignat 1856BR05, CaecilioidesBR05; Caeciliae, Caecilianellea, Cecilioididae, CecilioidinaeBR17]

*Cecilioides acicula (Müller 1774)BR17 [=Buccinum aciculaBR17, Achatina (Acicula) aciculaG40, Bulimus aciculaG40, Cionella aciculaG40; incl. Helix octonaG40, Buccinum terrestreG40]

Eremopeas tuckeri (Pfeiffer 1846)SS10 [=Bulimus tuckeriS88, Prosopeas (Eremopeas) tuckeriS88; incl. B. walli Cox 1864SS10]

Geostilbia gundlachi (Pfeiffer 1850) [incl. Cecilioides aperta (Swainson 1840) (n. n.), *G. caledonica Crosse 1867]SS10

*Opeas goodallii (Miller 1822)BR17 [=Bulimus goodalliiBR17, Helix goodalli non Ferussac 1821S88; incl. H. cochlicella clavulus (n. n.)G40, Achatina clavulusG40, Bulimus clavulusG40, B. pumilus Pfeiffer 1840S88, Opeas pumilumS88]

Prosopeas achatinaceum (Pfeiffer 1846) [=Bulimus achatinaceus; incl. Achatina javanica Reeve 1849, Opeas javanicum]S88

*Type species of generic name indicated

References

[A66] Adams, H. 1866. List of land and freshwater shells collected by Mr. E. Bartlett on the Upper Amazons, and on the River Ucayali, eastern Peru, with descriptions of new species. Proceedings of the Zoological Society of London 1866: 440–445.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[CC06] Cameron, R. A. D., L. M. Cook, G. A. Goodfriend & M. B. Seddon. 2006. Fossil land snails of Porto Santo, Madeiran archipelago: change and stasis in Pleistocene to Recent times. Malacologia 49 (1): 25–59.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[D66] Dohrn, H. 1866. Synopsis of the birds of Ilha do Principe, with some remarks on their habits and descriptions of new species. Proceedings of the Zoological Society of London 1866: 324–332.

[E99] Esu, D. 1999. Contribution to the knowledge of Neogene climate changes in western and central Europe by means of non-marine molluscs. In: J. Agustí, L. Rook & P. Andrews (eds) The Evolution of Neogene Terrestrial Ecosystems in Europe pp. 328–354. Cambridge University Press.

[FGN07] Fontaine, B., O. Gargominy & E. Neubert. 2007. Land snail diversity of the savanna/forest mosaic in Lopé National Park, Gabon. Malacologia 49 (2): 313–338.

[G40] Gray, J. E. 1840. A Manual of the Land and Fresh-water Shells of the British Islands, with figures of each of the kinds. By William Turton, M.D. A new edition, thoroughly revised and much enlarged. Longman, Orme, Brown, Green, and Longmans: London.

[J68] Jacobson, M. K. 1968. On a collection of terrestrial mollusks from Nicaragua. Nautilus 81 (4): 114–119.

[NB19] Nardi, G., A. Braccia, S. Cianfanelli & M. Bodon. 2019. Revision of the systematic position of Lindbergia garganoensis Gittenberger & Eikenboom, 2006, with reassignment to Vitrea Fitzinger, 1833 (Gastropoda, Eupulmonata, Pristilomatidae). Basteria 83 (1–3): 19–28.

[P02] Pickford, M. 2002. Early Miocene grassland ecosystem at Bukwa, Mount Elgon, Uganda. Comptes Rendus Palevol 1: 213–219.

[PB27] Pilsbry, H. A., & J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of Congo malacology. Bulletin of the American Museum of Natural History 53 (2): 69–602, pls 10–77.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[TC89] Tapparone Canefri, T. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XVIII.—Molluschi terrestri e d’acqua dolce. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 295–359.

[T68] Thompson, F. G. 1968. Two noteworthy achatinids from Panama. Nautilus 81 (3): 105.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[W79] Walker, B. 1879. Catalogue of the shell-bearing Mollusca of Michigan. Journal of Conchology 2: 325–337.

Leave a comment

Your email address will not be published. Required fields are marked *