Rosy bitterling Rhodeus ocellatus ocellatus, copyright Seotaro.

Belongs within: Cypriniformes.

The Acheilognathinae, bitterlings, are a group of cyprinid fishes found in Europe and eastern Asia, in which females develop a long ovipositor when spawning which they commonly use to deposit eggs in the mantle cavity of molluscs (Bond 1996).

    |–Tanakia B96
    |    |–T. lanceolata T90
    |    |–T. limbata T90
    |    `–T. tanago (Tanaka 1909) I92
    |–Acheilognathus B96
    |    |–A. cyanostigma T90
    |    |–A. longipinnis Regan 1905 I92
    |    |–A. melanogaster T90
    |    |–A. rhombeus T90
    |    |–A. tabira T90
    |    `–A. typus (Bleeker 1863) I92
    `–Rhodeus B96
         |–R. atremius I92
         |    |–R. a. atremius I92
         |    `–R. a. suigensis (Mori 1935) I92
         |–R. ocellatus I92
         |    |–R. o. ocellatus I92
         |    |–R. o. kurumeus Jordan & Thompson 1914 I92
         |    `–R. o. smithii T90
         `–R. sericeus (Pallas 1776) SE08
              |–R. s. sericeus CS77
              `–R. s. amarus CS77 [=Cyprinus amarus D56]

*Type species of generic name indicated


[B96] Bond, C. E. 1996. Biology of Fishes 2nd ed. Saunders College Publishing: Fort Worth.

[CS77] Cramp, S., & K. E. L. Simmons (eds) 1977. Handbook of the Birds of Europe, the Middle East and North Africa: The Birds of the Western Palaearctic vol. 1. Ostrich to Ducks. Oxford University Press: Oxford.

[D56] Dawes, B. 1956. The Trematoda with special reference to British and other European forms. University Press: Cambridge.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[SE08] Sevcsik, A., & T. Erös. 2008. A revised catalogue of freshwater fishes of Hungary and the neighbouring countries in the Hungarian Natural History Museum (Pisces). Annales Historico-Naturales Musei Nationalis Hungarici 100: 331–383.

[T90] Taguchi, S. 1990. Nihon no Sakana. Kogakukan: Tokyo.

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