Achipteria coleoptrata, from here.

Belongs within: Poronoticae.

Achipteria is a genus of oribatid mites found in Eurasia and the Americas with long humeral processes and a sacculate octotaxic system. Two subgenera are recognised with species of Achipteria sensu stricto having long interlamellar setae that project considerably beyond the bases of the cuspides whereas in Izuachipteria they are minute or absent (Balogh & Balogh 1992).

Characters (from Balogh & Balogh 1992): Prodorsum covered by two large lamellae; interlamellar setae arising aligned with dorsosejugal suture or absent. Notogaster with ten pairs of setae; four pairs of sacculi; long, pointed horizontal process on shoulder; pteromorphae immovable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae; Legs tridactyle.

<==Achipteria Berlese 1885S04
    |  i. s.: A. bellusG46
    |         A. obscura Krivolutsky & Ryabinin 1976S93
    |         A. magnusG45
    |         A. punctataG45
    |--A. (Achipteria)S04
    |    |--A. (A.) coleoptrata (Linnaeus 1758)S04 [=Acarus coleoptratusS04, Notaspis coleoptrataBA85]
    |    |    |--A. c. coleoptrataS04 (see below for synonymy)
    |    |    `--A. c. major Mihelčič 1963S04
    |    |--A. (A.) acuta Berlese 1908S04 (see below for synonymy)
    |    |--A. (A.) armata (Banks 1895) [=Oribatella armata]S04
    |    |    |--A. a. armataS04
    |    |    `--A. a. spinosa Jacot 1938S04 [=A. armatum spinosumJ38]
    |    |--A. (A.) bentonensis Marshall, Reeves & Norton 1987 (see below for synonymy)S04
    |    |--A. (A.) bicarinata Moskacheva 1973S04
    |    |--A. (A.) borealis (Banks 1899) (n. d.) [=Oribatella borealis]S04
    |    |--A. (A.) catskillensis Nevin 1977S04
    |    |--A. (A.) clarencei Nevin 1977S04
    |    |--A. (A.) cucullata Moskacheva 1973S04
    |    |--A. (A.) curta Aoki 1970S04
    |    |--A. (A.) dubia Weigmann 2001S04
    |    |--A. (A.) elegans Schweizer 1956S04
    |    |--A. (A.) hasticeps (Hull 1914) (n. d.)S04 [=Oribata hasticepsS04, Notaspis hasticepsL87]
    |    |--A. (A.) holomonensis Cancela da Fonseca & Stamou 1987S04
    |    |--A. (A.) italica (Oudemans 1913) [=Notaspis italica]S04
    |    |--A. (A.) languida (Nicolet 1855) (n. d.) [=Oribata languida]S04
    |    |--A. (A.) longesensillus Schweizer 1956S04
    |    |--A. (A.) mayeriana Palacios-Vargas & Socarrás 2001S04
    |    |--A. (A.) minuta (Ewing 1909) (n. d.) [=Notaspis minuta]S04
    |    |--A. (A.) moderatior Berlese 1923 (n. d.) [=A. nitens moderatior]S04
    |    |--A. (A.) nuda (Hall 1911) (n. d.) [=Notaspis nuda]S04
    |    |--A. (A.) oregonensis Ewing 1918S04
    |    |--A. (A.) oudemansi Jacot 1929 (n. d.)S04
    |    |--A. (A.) praeoccupata Subías 2004 [=A. oudemansi Hammen 1952 non Jacot 1929]S04
    |    |--A. (A.) ‘quadridentata’ (Willmann 1951) [=Notaspis quadridentata non N. bilipis quadridentata Haller 1882]S04
    |    |--A. (A.) regalis Berlese 1908S04
    |    |--A. (A.) sellnicki Hammen 1952S04
    |    |--A. (A.) serrata Hirauchi & Aoki 1997S04
    |    |--A. (A.) setulosa Golosova 1981S04
    |    |--A. (A.) sumatrensis Willmann 1931S04
    |    `--A. (A.) verrucosa Rjabinin 1974S04
    `--A. (Izuachipteria Balogh & Mahunka 1979) [incl. Hokkachipteria Balogh & Mahunka 1979]S04
         |--A. (*I.) imperfecta Suzuki 1972S04
         `--A. (I.) alpestris Aoki 1973S04 [=A. (*Hokkachipteria) alpestrisBM79]

Achipteria (Achipteria) acuta Berlese 1908S04 [incl. Notaspis anglicus Oudemans 1914S04, N. mixtus Sellnick 1943S04, Achipteria nitens Kishida 1930 non Oribata nitens Nicolet 1855S04, Oribata nitens Nicolet 1855 nec Gervais 1844 nec Johnston 1853S04, A. ovalis var. nitensL87, Oribates ovalis var. nitensL87]

Achipteria (Achipteria) bentonensis Marshall, Reeves & Norton 1987 [incl. A. oregonensis borealis Ewing 1918 non Oribatella borealis Banks 1899]S04

Achipteria (Achipteria) coleoptrata coleoptrata (Linnaeus 1758)S04 [incl. Oribata intermedia Michael 1898S04, Oribates nicoletii Berlese 1883S04, Oribata nicoletiS04, *Achipteria nicoletiS04, Parachipteria nicoletiS07, Oribata ovalis Nicolet 1855 non Koch 1835S04, Achipteria ovalisL87, Oribates ovalisB83, Oribata redux Hull 1914S04, A. reduxL87]

*Type species of generic name indicated


[BM79] Balogh, J., & S. Mahunka. 1979. New data to the knowledge of the oribatid fauna of the Neogea (Acari). IV. Acta Zoologica Academiae Scientiarum Hungaricae 25 (1–2): 35–60.

[B83] Berlese, A. 1883. Acari, Myriopoda et Scorpiones hucusque in Italia reperta. Fascicle 3. Typis Fratelli Salmin: Patavii.

[BA85] Bernini, F., & R. Arcidiacono. 1985. Notulae oribatologicae XXXVI. A catalogue of the Sicilian oribatids (Acarida, Oribatida). Animalia 12 (1–3): 165–205.

[G45] Grandjean, F. 1945. La signification évolutive de quelques caractères des acariens (1re série). Bulletin Biologique de la France et de la Belgique 79 (4): 297–325.

[G46] Grandjean, F. 1946. Les poils et les organes sensitifs portés par les pattes et le palpe chez les oribates. Troisième partie. Bulletin de la Société Zoologique de France 71: 10–29.

[J38] Jacot, A. P. 1938. Some new western North Carolina moss-mites. Proceedings of the Entomological Society of Washington 40 (1): 10–15.

[L87] Luxton, M. 1987. The oribatid mites (Acari: Cryptostigmata) of J. E. Hull. Journal of Natural History 21: 1273–1291.

[S07] Schatz, H. 2007. Biogeography of oribatid mites (Acari, Oribatida) from the Cordillera de Talamanca, Costa Rica and Panama. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 151–167. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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