Acochlidimorpha

Unela remanei, from Arnaud et al. (1986). Length = 3 mm.

Belongs within: Heterobranchia.

The Acochlidimorpha are a group of small, mostly aquatic slugs that usually bear a large visceral hump projecting freely from the headfoot.

<==Acochlidimorpha (see below for synonymy)
|–Acochlidioidea [Strubellioidea, Strubellioidei] BR17
|    |–Pseudunela Salvini-Plawen 1973 [Pseudunelidae, Pseudunelloidea] BR05
|    |    `–*P. cornuta (Challis 1970) [=Microhedyle cornuta] BR17
|    |–Aiteng Swennen & Buatip 2009 [Aitengidae] BR17
|    |    `–*A. ater Swennen & Buatip 2009 BR17
|    |–Bathyhedyle Neusser, Jörger et al. 2016 [Bathyhedylidae] BR17
|    |    `–*B. boucheti Neusser, Jörger et al. 2016 BR17
|    |–Hedylopsis Thiele 1931 BR05 [Hedylopsidae BR17, Hedylopsinae]
|    |    `–*H. spiculifera (Kowalewsky 1901) BR17 (see below for synonymy)
|    |–Tantulum Rankin 1979 BR05 [Pharyngoneura, Tantulidae BR17, Tantuloidea]
|    |    `–*T. elegans Rankin 1979 BR17
|    `–Acochlidiidae BR17
|         |–Acochlidium Strubell 1892 [Acochlidiinae] BR05
|         |    `–*A. amboinense Strubell 1892 BR17
|         |–Palliohedyle Rankin 1979 BR17, BR05 (see below for synonymy)
|         |    `–*P. weberi (Bergh 1895) [=*Hedyle weberi] BR17
|         `–Strubellia Odhner 1937 BR17, BR05 [Strubelliidae]
|              `–*S. paradoxa (Strubell 1892) [=Acochlidium paradoxum] BR17
`–Parhedyloidea BR17
|–Asperspinidae BR17
|    |–Minicheviella Starobogatov 1983 BR17, BR05 [Minicheviellidae, Minichevielloidea]
|    |    `–*M. murmanica (Kudinskaja & Minichev 1978) [=Hedylopsis murmanica] BR17
|    `–Asperspina Rankin 1979 BR05
|         |–*A. brambelli (Swedmark 1968) [=Hedylopsis brambelli] BR17
|         |–A. loricata (Swedmark 1968) APS-P86
|         |–A. murmanica (Kudinskaja & Minichev 1978) APS-P86
|         |–A. rhopalotecta (Salvini-Plawen 1973) APS-P86
|         `–A. riseri (Morse 1976) APS-P86
`–Parhedylidae [Avelariacea, Ganitidae, Ganitoidea, Microhedylidae, Microhedylinae, Velariacea] BR05
|–Parhedyle Thiele 1931 [Parhedylinae] BR05
|    `–*P. tyrtowii (Kowalewsky 1901) BR17 [=Hedyle tyrtowii BR17, Microhedyle tyrtowii APS-P86]
|–Sabulincola Rankin 1979 [Sabulincolidae] BR05
|    `–*S. odhneri (Marcus & Marcus 1955) [=Unela odhneri] BR17
|–Ganitus Marcus 1953 BR17, BR05
|    `–*G. evelinae Marcus 1953 BR17
|–Paraganitus Challis 1968 APS-P86
|    `–P. ellynnae Challis 1968 APS-P86
|–Livorniella Rankin 1979 BR17, BR05 [Livorniellidae, Livornielloidea, Visceroneura]
|    `–*L. glomerans (Salvini-Plawen 1973) [=Microhedyle glomerans] BR17
|–Pontohedyle Golikov & Starobogatov 1972 BR05 (see below for synonymy)
|    |–*P. milaschewitchii (Kowalewsky 1901) [=Hedyle milaschewitchii, *Mancohedyle milaschewitchii] BR17
|    |–P. brasilensis (Rankin 1979) APS-P86
|    `–P. verrucosa (Challis 1970) APS-P86
|–Microhedyle Hertling 1930 BR05
|    |–*M. lactea Hertling 1930 BR17
|    |–M. cryptophthalma Westheide & Wawra 1974 APS-P86
|    |–M. gerlachi Marcus & Marcus 1959 APS-P86
|    `–M. milaschewitchii M62
`–Unela Marcus 1953 [Unelidae] BR05
|–*U. remanei Marcus 1953 BR17
|–U. glandulifera (Kowalevsky 1901) APS-P86
|–U. glomerans (Salvini-Plawen 1973) APS-P86
|–U. nahatensis Doe 1974 APS-P86
`–U. neapolitana (Rankin 1979) [=Stellaspina neapolitana] APS-P86

Nomina nuda: Microhedyle fundensis Rankin in Arnaud, Poizat & Salvini-Plawen 1986 APS-P86
Pseudunela eirenei Wawra in Arnaud, Poizat & Salvini-Plawen 1986 APS-P86

Acochlidimorpha [Acochlidiacea, Acochlidiida, Acochlidiomorpha, Cerebroneura, Euacochlidiacea, Euacochlidioidea, Hedylopsoidea, Hedylopsoidei, Microhedylacea, Pedoneura]

*Hedylopsis spiculifera (Kowalewsky 1901) BR17 [=Hedyle spiculifera BR17; incl. Hedylopsis suecica Odhner 1937 APS-P86]

Palliohedyle Rankin 1979 BR17, BR05 [=Hedyle Bergh 1895 nec Guenée 1857 nec Malmgren 1865 BR05; Hedylinae, Palliohedylidae, Palliohedyloidea, Palliohedyloidei]

Pontohedyle Golikov & Starobogatov 1972 [incl. Mancohedyle Rankin 1979 APS-P86, BR05; Mancohedylidae, Pontohedylidae]

*Type species of generic name indicated

REFERENCES

[APS-P86] Arnaud, P. M., Cl. Poizat & L. v. Salvini-Plawen. 1986. Marine-interstitial Gastropoda (including one freshwater interstitial species).  In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 153–176. E. J. Brill/Dr W. Backhuys: Leiden.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

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