Acteon punctocoelata, copyright J. Goddard.

Belongs within: Acteonimorpha.
Contains: Pupa.

The Acteonidae are a group of marine gastropods of uncertain affinities. Though anatomically similar to opisthobranchs, they possess certain plesiomorphic features such as the presence of an operculum to close the shell aperture. The shell is well calcified with a distinct spire and an elongate aperture. The radula usually lacks a rachidian tooth and possesses from five to over two hundred laterals each side.

<==Acteonidae [Pupacea, Pupidae, Tornatelladae, Tornatellidae, Tornatellinae]
    |--Liocarenus Harris & Burrows 1891 [Liocareninae]BR05
    |    `--*L. conovuliformis (Deshayes 1824) [=Auricula conovuliformis]BR17
    `--Acteoninae [Actaeoninae]BR05
         |--Eoactaeon Stephenson 1955KB01
         |--Nucleopsis Conrad 1865 [Nucleopsinae]BR05
         |    `--*N. subvaricata (Conrad 1860) [=Acteonina subvaricata]BR17
         |--Tornatellaea Conrad 1860 [Tornatellaeinae]BR05
         |    |--*T. bella Conrad 1860BR17
         |    |--T. cretacea Wade 1926KB01
         |    |--T. fontis (Duméril 1806)TTE93
         |    |--T. globulosa Wade 1926KB01
         |    `--T. hebertiTTE93
         |--Tornatella Lamarck 1816BR05
         |    |--*T. flammea (Bruguière 1789)BR17 (see below for synonymy)
         |    |--T. bullataR42
         |    |--T. glabra Reeve 1842R42
         |    |--T. minuta Petterd 1879P79
         |    |--T. oryza Reeve 1842R42
         |    |--T. ovalisH86
         |    |--T. siebaldii Reeve 1842R42
         |    |--T. tessellata Reeve 1842R42
         |    |--T. venustaR42
         |    `--T. virgata Reeve 1842R42
         `--Acteon Montfort 1810 [=Actaeon (l. c.) non Rang 1829]BR05
              |  i. s.: A. affinis (Sowerby 1836) [=Tornatella affinis]F27a
              |         A. ambiguus (Hutton 1885) [=Admete ambigua]PH90
              |         ‘Actaeon’ articulatusD56
              |         A. chattonensisPB29
              |         A. cicatricosus Sohl 1964KBP02
              |         A. cratericulatus Hedley 1906P61 [=A. craticulatus Murdoch & Suter 1906F27a]
              |         ‘Actaeon’ muelleri Bosquet 1861KB01
              |         A. oneroaensis Powell & Bartrum 1929PB29
              |         A. ooliticusKBP02
              |         A. panamensis Dall 1908PP78
              |         A. praestitus Finlay 1924 [=A. sulcatus (Hutton 1885) (preoc.)]F27b
              |         A. roseus Hedley 1906F27a
              |         A. tenellusN79
              |         A. traski Stearns 1898O27
              |--A. (Acteon)O27
              |    |--*A. (A.) tornatilis (Linnaeus 1758)BR17 (see below for synonymy)
              |    `--A. (A.) fasciatusO27 [=Tornatella fasciataO27; incl. Auricula bifasciataR42, T. bifasciataG20]
              `--A. (Rictaxis Dall 1871)O27
                   |--A. (R.) painei Dall 1903O27
                   `--A. (R.) punctocoelata (Carpenter 1864) [=Tornatella punctocoelata]O27
                        |--A. p. punctocoelataO27
                        `--A. p. vancouverensis Oldroyd 1927O27
Acteonidae incertae sedis:
  Dactylus Schumacher 1817BR05
    |--D. maurusF66
    `--D. mustelinusF66
  Microglyphis Dall 1902O27
    |--*M. curtulus [=Acteon curtulus]O27
    |--M. breviculus Dall 1902O27
    |--M. estuarinus Dall 1908O27
    `--M. globularis (Smith 1875) [=Admete globularis]PH90
  Bullinula Swainson 1840P61
    `--*B. lineata (Gray 1825)P61 (see below for synonymy)
  Neactaeonina Thiele 1912P61
    |--*N. cingulata [=Acteonina cingulata]P61
    `--N. inexpectata Dell 1956P61
  Crenilabium Cossmann 1889BS00
    `--C. (Eoacteon Stephenson 1955)BS00
         |--C. (*E.) linteus (Conrad 1858) [=Solidulus linteus, Actaeon linteus]BS00
         `--C. (E.) valdovinosensis Bandel & Stinnesbeck 2000BS00
  Japonacteon suturalis (Adams 1855)WBS-S15

*Acteon (Acteon) tornatilis (Linnaeus 1758)BR17 [=Bulla tornatilisBR17, Bulimus tornatilisR42, Voluta tornatilisKBP02]

*Bullinula lineata (Gray 1825)P61 [=Bulla lineataP61, Bullina lineataBR17; incl. Bulla scabra Gmelin 1791 (preoc.)BR17, F27a, Bullina scabraF27a, Voluta ziczac Muhlfeldt 1878 non Schroeter 1804F27a]

*Tornatella flammea (Bruguière 1789)BR17 [=Buccinum flammeumBR17, Actaeon flammeusH09, Voluta flammeaR42; incl. Bulimus variegatusR42]

*Type species of generic name indicated


[BS00] Bandel, K., & W. Stinnesbeck. 2000. Gastropods of the Quiriquina Formation (Maastrichtian) in central Chile: paleobiogeographic relationships and the description of a few new taxa. Zbl. Geol. Paläont. Teil I 7–8: 757–788.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[D56] Dell, R. K. 1956. The archibenthal Mollusca of New Zealand. Dominion Museum Bulletin 18: 1–235.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[F66] Fraser, L. 1866. Communication of a list of mollusks collected by R. Swinhoe, Esq., in Formosa. Proceedings of the Zoological Society of London 1866: 146.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H86] Hutton, F. W. 1886. The Mollusca of the Pareora and Oamaru systems of New Zealand. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 205–237.

[KB01] Kiel, S., & K. Bandel. 2001. About Heterostropha (Gastropoda) of the Campanian of Torallola, Spain. Journal of the Czech Geological Society 46 (3–4): 319–334.

[KBP02] Kiel, S., K. Bandel & M. del C. Perrilliat. 2002. New gastropods from the Maastrichtian of the Mexcala Formation in Guerrero, southern Mexico, part II: Archaeogastropoda, Neritimorpha and Heterostropha. Neues Jahrbuch für Geologie und Paläontologie—Abhandlungen 226 (3): 319–342.

[N79] Norman, A. M. 1879. The Mollusca of the fiords near Bergen, Norway. Journal of Conchology 2: 8–77.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 1. Stanford University Press: Stanford University (California).

[PH90] Petit, R. E. & M. G. Harasewych. 1990. Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia). Nautilus 103 (Suppl. 1): 1–69.

[P79] Petterd, W. F. 1879. New species of Tasmanian marine shells. Journal of Conchology 2: 102–105.

[PP78] Poorman, F. L., & L. H. Poorman. 1978. Additional molluscan records from Bahía de Los Angeles, Baja California Norte. Veliger 20 (4): 369–374.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[PB29] Powell, A. W. B., & J. A. Bartrum. 1929. The Tertiary (Waitematan) molluscan fauna of Oneroa, Waiheke Island. Transactions and Proceedings of the New Zealand Institute 60: 395–447.

[R42] Reeve, L. 1842. Monograph of the genus Tornatella. Proceedings of the Zoological Society of London 10: 58–62.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[WBS-S15] Willan, R. C., C. Bryce & S. M. Slack-Smith. 2015. Kimberley marine biota. Historical data: molluscs. Records of the Western Australian Museum Supplement 84: 287–343.

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