Aderidae

 Megaxenus termitophilus, from Hangay & Zborowski (2010).

Belongs within: Tenebrionoidea.

The Aderidae are a group of small, often colourful beetles typically found on vegetation, often resting on the undersides of leaves. Larvae have been collected in rotting wood, under bark or in leaf litter, or in insect nests (Chandler 2002).

Characters (from Lawrence & Britton 1991): Elongate, convex to somewhat flattened beetles; elytra commonly bicoloured, vestiture consisting of erect and/or decumbent hairs, the latter sometimes forming a pattern; head abruptly constricted behind eyes; eyes relatively large, coarsely faceted; antennae 11-segmented, filiform or flabellate; maxillary palps securiform; pronotum narrow, lacking lateral carinae; fore coxae projecting, trochantins concealed, fore coxal cavities externally and internally open; mid coxal cavities narrowly separated, laterally open; tarsal formula 5-5-4, antepenultimate tarsal segment lobed, penultimate one reduced; abdomen with five ventrites, the first two ventrites strongly connate or solidly fused (in which case abdomen appears to have four ventrites); hind femora of males sometimes bearing setose secretory organs. Larvae almost always narrowly elongate, parallel sided, somewhat flattened and lightly sclerotised, except for tips of urogomphi. Mandible almost always with hyaline lobe at base of mola, median endocarina on head and armature on S9 absent.

<==Aderidae [Circaeidae, Euglenesidae, Euglenidae, Hylophilidae, Xylophilidae]
    |  i. s.: Syzeton abnormisLB91
    |         Pseudananca ruficollisLB91
    |         SyzetonellusLB91
    |         CircaeusP92
    |         MegaxenusB14
    |           |--M. bioculatusB14
    |           |--M. papuensisB14
    |           `--M. termitophilus Lawrence 1990B14
    |         EscalerosiaC02
    |--Emelinus Casey 1895 [Emelinini]C02
    |    `--E. butleriC02
    |--EuglenesiniC02
    |    |--PseudolotelinaC02
    |    `--Zonantes Casey 1895 [Euglenesina]C02
    |--PhytobaeniniC02
    |    |--Axylophilus Casey 1895C02
    |    |    `--A. yuccaeC02
    |    |--Ganascus Casey 1895 [incl. Sandytes Casey 1895]C02
    |    |    |--G. ptinoidesC02
    |    |    `--G. ventricosusC02
    |    `--PhytobaenusRS15
    `--AderiniC02
         |--Gymnoganascus Werner 1990 [Olotelina]C02
         |    `--G. stephaniC02
         |--Cnopus Champion 1893 [Cnopina]C02
         |    |--C. impressusC02
         |    `--C. nucleusC02
         |--Aderus Stephens 1829 (see below for synonymy)C02
         |    |--‘Xylophilus’ brouni Pic 1901E02
         |    |--A. brunnipennisC02
         |    |--‘Xylophilus’ constrictusC02
         |    |--A. populneusC02
         |    `--‘Xylophilus’ undatus Gemming 1870 [incl. X. fasciatus Bohem. 1858]M86
         `--SyzetonininaC02
              |--SyzetoninusLB91
              |--Elonus Casey 1895C02
              |--Vanonus Casey 1895 [incl. Tanilotes Casey 1895]C02
              |    `--V. densusC02
              |--Ariotus Casey 1895 [incl. Scanylus Casey 1895]C02
              |    `--A. quercicolaC02
              `--Pseudariotus Casey 1895C02
                   `--P. notatusC02

Aderus Stephens 1829 [incl. Hylophilus Berthold 1827 non Temminck 1822, Phomalus Casey 1895, Xylophilus Latreille 1825 non Mannerheim 1823; Aderina]C02

*Type species of generic name indicated

References

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[C02] Chandler, D. S. 2002. Aderidae Winkler 1927. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 559–563. CRC Press.

[E02] Emberson, R. M. 2002. The beetle (Coleoptera) fauna of the Chatham Islands: additions and corrections. New Zealand Entomologist 25: 69–77.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[RS15] Robertson, J. A., A. Ślipiński, M. Moulton, F. W. Shockley, A. Giorgi, N. P. Lord, D. D. McKenna, W. Tomaszewska, J. Forrester, K. B. Miller, M. F. Whiting & J. V. McHugh. 2015. Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia). Systematic Entomology 40: 745–778.

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