Lateral (above), dorsal (below left) and ventral views of Epieremulus sp., from Norton & Behan-Pelletier (2009).

Belongs within: Holonota.

The Anderemaeidae are a group of oribatid mites found in temperate regions and at higher altitudes in the tropics.

Characters (from Norton & Behan-Pelletier 2009): Length 280–850 µm. Gnathosoma normal; chelicerae chelate-dentate, with two setae; fixed digit of chelicerae with unsclerotised lobe with 7–10 small teeth. Prodorsum with costulae, without true lamellae; bothridium with internal spiral thickening; tutorium, humeral apophysis, parastigmatic enantiophysis S and enantiophysis V present. Notogaster oval; octotaxic system absent. Pedotectum II present. Tarsus II with two solenidia.

    |--Cristeremaeus Balogh & Csiszár 1963NB-P09, S04
    |    |--*C. humeratus Balogh & Csiszár 1963S04
    |    `--C. clavatus Mahunka 1980S04
    |--Yungaseremaeus Balogh & Mahunka 1969NB-P09, S04
    |    `--*Y. longisetosus Balogh & Mahunka 1969S04
    |--Anderemaeus Hammer 1958S04
    |    |--*A. monticola Hammer 1958S04
    |    |--A. australiensis Balogh & Balogh 1983S04
    |    |--A. capitatus Balogh & Balogh 1985S04
    |    |--A. chilensis Hammer 1962S04
    |    |--A. hammerae Mahunka 1980S04
    |    |--A. hidasii Balogh 1995S04
    |    |--A. magellanis Hammer 1962S04
    |    `--A. sturmi Balogh & Balogh 1985S04
    `--Epieremulus Berlese 1916PL17, S04 [incl. Carabodoides Jacot 1937S04]
         |--*E. geometricus (Berlese 1916)S04 (see below for synonymy)
         |--E. apicalis (Banks 1895) [=Carabodes apicalis]S04
         |--E. braziliensis (Balogh & Mahunka 1969) [=Carabodoides braziliensis]S04
         |--E. circulus Yamamoto & Aoki 2000S04
         |--E. frontatus (Warburton 1912) [=Notaspis frontata]S04
         |--E. granulatus (Balogh & Mahunka 1979) [=Carabodoides granulatus]S04
         |    |--E. g. granulatusS04
         |    `--E. g. andicola (Balogh 1988) [=Carabodoides granulatus andicola]S04
         |--E. humeratus (Aoki 1987) [=Carabodoides humeratus]S04
         |--E. laticeps Balogh 1963S04 [=Carabodoides laticepsBB85]
         |--E. longicarinatus (Balogh & Mahunka 1978) [=Carabodoides longicarinatus]S04
         `--E. longiseta (Balogh 1988) [=Carabodoides longiseta]S04

*Epieremulus geometricus (Berlese 1916)S04 [=Eremulus (*Epieremulus) geometricusS04; incl. *Carabodoides saccharomycetoides Jacot 1937B65, S04]

*Type species of generic name indicated


[B65] Balogh, J. 1965. A synopsis of the world oribatid (Acari) genera. Acta Zoologica Academiae Scientiarum Hungaricae 11 (1–2): 5–99.

[BB85] Balogh, J., & P. Balogh. 1985. Studies on the Anderemaeidae J. Balogh, 1972 (Acari, Oribatei). Opuscula Zoologica Budapest 19–20: 41–48.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[PL17] Pachl, P., A. C. Lindl, A. Krause, S. Scheu, I. Schaefer & M. Maraun. 2017. The tropics as an ancient cradle of oribatid mite diversity. Acarologia 57 (2): 309–322.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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