Ankylosaurinae

Skeletal reconstruction of Pinacosaurus, copyright Kabacchi.

Belongs within: Ankylosauridae.

The Ankylosaurinae is a clade of ankylosaurs from the Late Cretaceous of Asia and North America that has been defined as including all taxa closer to Ankylosaurus than Shamosaurus (Vickaryous et al. 2004). Well known members of the group include the North American Euoplocephalus tutus, one of the the largest ankylosaurs reaching a length of around six metres.

Synapomorphies (from Vickaryous et al. 2004): Nuchal shelf obscuring occiput in dorsal view; quadrate condyle obscured laterally by quadratojugal boss; buccal emargination strongly concave.

AnkylosaurinaeVMW04
|--Tsagantegia Tumanova 1993VMW04
| `--T. longicranialis Tumanova 1993VMW04
`--+--Tarchia Maryańska 1977VMW04
| `--T. gigantea (Maleev 1956) [=Dyoplosaurus giganteus; incl. T. kielanae Maryańska 1977]VMW04
`--+--+--Euoplocephalus Lambe 1910 (see below for synonymy)VMW04
| | `--E. tutus (Lambe 1902)VMW04 (see below for synonymy)
| `--Ankylosaurus Brown 1908VMW04
| `--A. magniventris Brown 1908VMW04
`--+--Saichania Maryańska 1977VMW04
| `--S. chulsanensis Maryańska 1977VMW04
|--Talarurus Maleev 1952VMW04
| |--T. disparoserratus Maleev 1952C01
| `--T. plicatospineus Maleev 1952VMW04
`--+--Nodocephalosaurus Sullivan 1999C01, VMW04
| `--N. kirtlandensis Sullivan 1999VMW04
`--Pinacosaurus Gilmore 1933VMW04 [incl. NinghsiasaurusN85, VirminicaudusN85]
|--P. grangeri Gilmore 1933 [incl. P. ninghsiensis Young 1935, Syrmosaurus viminicaudus Maleev 1952]VMW04
`--+--P. mephistocephalus Godefroit, Pereda-Suberbiola & Dong 1999VMW04
`--Tianzhenosaurus Pang & Cheng 1998VMW04
`--T. youngi Pang & Cheng 1998VMW04

Euoplocephalus Lambe 1910 [=Stereocephalus Lambe 1902 (preoc.); incl. Anodontosaurus Sternberg 1929, Dyoplosaurus Parks 1924, Scolosaurus Nopcsa 1928]VMW04

Euoplocephalus tutus (Lambe 1902)VMW04 [=Stereocephalus tutusVMW04; incl. Dyoplosaurus acutosquameus Parks 1924VMW04, Euoplocephalus acutosquameusD07, Scolosaurus cutleri Nopcsa 1929VMW04, E. cutleriN85, Anodontosaurus lambei Sternberg 1929VMW04]

*Type species of generic name indicated

References

[C01] Carpenter, K. 2001. Phylogenetic analysis of the Ankylosauria. In: Carpenter, K. (ed.) 2001. The Armored Dinosaurs pp. 455–483. Indiana University Press.

[D07] Dixon, D. 2007. The Complete Illustrated Encyclopedia of Dinosaurs & Prehistoric Creatures. Hermes House: London.

[N85] Norman, D. 1985. The Illustrated Encyclopedia of Dinosaurs. Salamander Books: London.

[VMW04] Vickaryous, M. K., T. Maryańska & D. B. Weishampel. 2004. Ankylosauria. In: Weishampel, D. B., P. Dodson & H. Osmólska (eds) The Dinosauria 2nd ed. pp. 363–392. University of California Press: Berkeley.

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