Cibicidoides wuellerstorfi, from Foram Barcoding.

Belongs within: Rotaliida.
Contains: Gavelinella, Heterolepa.

The Anomalinidae are a group of Foraminifera known from the Upper Triassic to the present (Loeblich & Tappan 1964b).

Characters (from Loeblich & Tappan 1964b): Test trochospiral to nearly planispiral, evolute on one or both sides; chambers simple; wall calcareous, coarsely perforate, granular in structure, bilamellar; primary aperture interiomarginal equatorial or somewhat extending onto spiral or umbilical sides, additional peripheral apertures may be present.

    |–Almaeninae LT64a
    |    |–Ganella Aurouze & Boulanger 1954 LT64b
    |    |    `–*G. neumannae Aurouze & Boulanger 1954 LT64b
    |    |–Queraltina Marie 1950 LT64b
    |    |    `–*Q. epistominoides Marie 1950 LT64b
    |    |–Anomalinella Cushman 1927 LT64b
    |    |    `–*A. rostrata (Brady 1881) [=Truncatulina rostrata] LT64b
    |    `–Almaena Samoylova 1940 (see below for synonymy) LT64b
    |         |–*A. taurica Samoylova 1940 LT64b
    |         |–*Kelyphistoma’ ampulloloculata Keijzer 1945 LT64b
    |         |–A. escornebovensis (Sigal 1949) [=*Planulinella escornebovensis] LT64b
    |         `–A. hieroglyphica Sigal 1950 [=A. (*Pseudoplanulinella) hieroglyphica] LT64b
    `–Anomalininae LT64a
         |–Gavelinella LT64b
         |–Angulogavelinella Hofker 1957 LT64b
         |    `–*A. gracilis (Marsson 1878) [=Discorbina gracilis] LT64b
         |–Asymmetrina Kristan-Tollmann 1960 LT64b
         |    `–*A. biomphalica Kristan-Tollmann 1960 LT64b
         |–Boldia van Bellen in van den Bold 1946 [=Terquemia van Bellen 1946 nec Tate 1868 nec Van Veen 1932] LT64b
         |    `–*B. lobata (Terquem 1882) [=Rotalina lobata, Anomalina lobata, *Terquemia lobata] LT64b
         |–Coleites Plummer 1934 LT64b
         |    `–*C. reticulosa (Plummer 1927) [=Pulvinulina reticulosa] LT64b
         |–Discanomalina Asano 1951 LT64b
         |    |–*D. japonica Asano 1951 LT64b
         |    `–D. semipunctata LJ98
         |–Holmanella Loeblich & Tappan 1962 LT64b
         |    `–*H. valmonteensis (Kleinpell 1938) [=Discorbinella valmonteensis] LT64b
         |–Involvina Kristan-Tollmann 1960 LT64b
         |    `–*I. obliqua Kristan-Tollmann 1960 LT64b
         |–Nummodiscorbis Hornibrook 1961 LT64b
         |    `–*N. novozealandica Hornibrook 1961 LT64b
         |–Paromalina Loeblich & Tappan 1957 LT64b
         |    `–*P. bilateralis Loeblich & Tappan 1957 LT64b
         |–Plagiostomella Kristan-Tollmann 1960 LT64b
         |    `–*P. inflata Kristan-Tollmann 1960 LT64b
         |–Pulsiphonina Brotzen 1948 LT64b
         |    `–*P. prima (Plummer 1927) [=Siphonina prima] LT64b
         |–Stensioina Brotzen 1936 LT64b
         |    |–*S. exsculpta (Reuss 1860) [=Rotalia exsculpta] LT64b
         |    `–S. clementiana G-B87
         |–Karreria Rzehak 1891 [incl. Vagocibicides Finlay 1939] LT64b
         |    |–*K. fallax Rzehak 1891 LT64b
         |    `–K. maoria (Finlay 1939) [=*Vagocibicides maoria] LT64b
         |–Anomalinoides Brotzen 1942 LT64b
         |    |–A. pinguis (Jennings 1936) [=Anomalina pinguis; incl. *Anomalinoides plummerae Brotzen 1942] LT64b
         |    |–A. globosus S05
         |    `–A. macraglabra H90
         |–Hanzawaia Asano 1944 LT64b
         |    |–*H. nipponica Asano 1944 LT64b
         |    |–H. concentrica JW99
         |    `–H. strettoni LT64a
         |–Anomalina d’Orbigny 1826 [incl. Aspidospira Ehrenberg 1844, Porospira Ehrenberg 1844] C40
         |    |–*A. punctulata d’Orbigny 1826 C40
         |    |–*Porospira’ comes Ehrenberg 1844 LT64b
         |    |–A. globulosa JW99
         |    |–A. grosserugosa C40
         |    |–A. henbesti M08
         |    `–A. miosuturalis B49
         |–Melonis de Montfort 1808 (see below for synonymy) LT64b
         |    |–M. pompilioides (Fichtel & Moll 1798) (see below for synonymy) LT64b
         |    |–M. affinis LJ98
         |    |–M. barleeanum JW99
         |    |–M. maorica H90
         |    |–M. nicobarensis (Cushman 1963) [=Nonion nicobarensis] H03
         |    |–‘Rotalia’ tuberculifera Reuss 1862 [=Gavelinonion tuberculiferum] LT64b
         |    |–‘Nautilus’ umbilicatulus Walker & Jacob in Kanmacher 1798 [=*Gavelinonion umbilicatulum] LT64b
         |    `–M. zaandami LT64b
         |–Cibicidoides Thalmann 1939 [incl. Parrelloides Hofker 1956] LT64b
         |    |–*C. mundula (Brady, Parker & Jones 1888) [=Truncatulina mundula] LT64b
         |    |–C. aknerianus (d’Orbigny 1846) [=Rotalina akneriana; incl. Cibicides asanoi Matsunaga 1963] H03
         |    |–C. hyalinus (Hofker 1951) [=Cibicides hyalinus, *Parrelloides hyalinus] LT64b
         |    |–C. involutus G-B87
         |    |–C. kullenbergi JW99
         |    |–C. mabahethi S05
         |    |–C. pseudoungerianus JW99
         |    |–C. robertsonianus JW99
         |    `–C. wuellerstorfi JW99
         `–Heterolepa LT64b

Almaena Samoylova 1940 [incl. Kelyphistoma Keijzer 1945, Planulinella Sigal 1949, Pseudoplanulinella Sigal 1950] LT64b

Melonis de Montfort 1808 [=Melonia bronn 1849 nec Lamarck 1822 nec Schinz 1825; incl. Gavelinonion Hofker 1957, Melossis Pallas in Oken 1815] LT64b

Melonis pompilioides (Fichtel & Moll 1798) [=Nautilus pompilioides, *Melossis pompilioides; incl. *Melonis etruscus de Montfort 1808] LT64b

*Type species of generic name indicated


[B49] Battey, M. H. 1949. The geology of the Tuakau-Mercer area, Auckland. Transactions and Proceedings of the Royal Society of New Zealand 77 (3): 429–455.

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3rd ed. Harvard University Press: Cambridge (Massachusetts).

[G-B87] Gawor-Biedowa, E. 1987. New benthic foraminifers from the Late Cretaceous of Poland. Acta Palaeontologica Polonica 32 (1–2): 49–72.

[H03] Hanagata, S. 2003. Miocene-Pliocene Foraminifera from the Niigata oil-fields region, northeastern Japan. Micropaleontology 49 (4): 293–340.

[H90] Hayward, B. W. 1990. Use of foraminiferal data in analysis of Taranaki Basin, New Zealand. Journal of Foraminiferal Research 20: 71–83.

[JW99] Jian, Z.-M., L.-J. Wang, M. Kienast, M. Sarnthein, W. Kuhnt, H.-L. Lin & P.-X. Wang. 1999. Benthic foraminiferal paleoceanography of the South China Sea over the last 40,000 years. Marine Geology 156: 159–186.

[LJ98] Li, Q., N. P. James, B. McGowran, Y. Bone & J. Cann. 1998. Synergetic influence of water masses and Kangaroo Island barrier on foraminiferal distribution, Lincoln and Lacepede shelves, South Australia: a synthesis. Alcheringa 22 (2): 153–176.

[LT64a] Loeblich, A. R., Jr & H. Tappan. 1964a. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[LT64b] Loeblich, A. R., Jr & H. Tappan. 1964b. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 2. The Geological Society of America and The University of Kansas Press.

[M08] McMillan, I. K. 2008. Reappraisal of foraminiferal assemblages of the Santonia-Campanian Mzamba Formation type section, and their correlation with the stratigraphic succession of the KwaZulu Basin. African Natural History 4: 25–34.

[S05] Semeniuk, T. A. 2005. Fossil foraminiferal assemblages from Pleistocene seagrass-bank deposits of the southern Perth Basin, Western Australia, and their palaeotemperature implications. Journal of the Royal Society of Western Australia 88 (4): 177–190.

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