Gymnarus auriculatus, from Stromboidea.de.

Belongs within: Stromboidea.

The Aporrhaidae, pelican’s feet, are a group of marine gastropods characterised by a shell with a large, laterally extended outer lip.

|--Harpagodes Gill 1870 [Harpagodesidae, Harpagodidae, Harpagodinae]BR05
| `--*H. pelagi (Brongniart 1821) [=Strombus pelagi]BR17
|--Spiniloma Gründel, Nutzel & Schulbert 2009BR17 (see below for synonymy)
| `--*S. longispina (Eudes-Deslongchamps 1843) [=Ranella longispina, *Spiniloma longispina]BR17
|--Dimorphosoma Gardner 1875 [Dimorphosominae]BR17
| `--*D. calcarata (Sowerby 1822) [=Rostellaria calcarata]BR17
|--Pterocerella Meek 1864 [Pterocerellidae, Pterocerellinae]BR17
| `--*P. tippana (Conrad 1858) [=Harpago tippana]BR17
|--Struthioptera Finlay & Marwick 1937BR05 [StruthiopterinaeBR17]
| |--*S. haastiana (Wilckens 1922) [=Arrhoges haastianus]S98
| `--S. novoseelandica (Wilckens 1922) [=Perissoptera novoseelandica]S98
|--Anchura Conrad 1860W77 [AnchurinaeBR17]
| |--*A. abrupta Conrad 1860BR17
| `--A. fusiformis Meek in White 1877W77
|--Arrhoginae [Perissopteridae, Perissopterinae]BR17
| |--Cerycium Philippi 1841 [incl. Arrhoges Gabb 1868]BR17
| | |--‘Drepanocheilus (Arrhoges)’ speciosus (Schlotheim 1820) [incl. *Cerycium paradoxum Philippi 1841]BR17
| | `--‘Rostellaria’ occidentalis Beck 1836 [=*Arrhoges occidentalis]BR17
| `--Perissoptera Tate 1865BR05
| |--*P. parkinsoni (Mantell 1822) [=Rostellaria parkinsoni]BR17
| `--P. thomsoni [=Hemichenopus thomsoni]A27
| |--‘Alaria’ Morris & Lycett 1851 nec Schrank 1788 nec Duncan 1841BR17, BR05 [Alariidae]
| | `--*A. armata Morris & Lycett 1851BR17
| |--Aporrhais de Costa 1778 [=Chenopus Philippi 1836; Chenopidae]BR05
| | |--*A. pespelecani (Linnaeus 1758) [=Strombus pespelecani, *Chenopus pespelecani]BR17
| | |--A. occidentalis Beck 1836S11
| | `--A. serresianus (Michaud 1828) (see below for synonymy)N79
| `--Dicroloma Gabb 1868BR17, BR05 [Dicrolomatidae, Dicrolomidae]
| |--*D. lorierei (d’Orbigny 1860) [=Pterocera lorierei]BR17
| |--‘Rostellaria’ bispinosa Phillips 1829 [=Alaria bispinosa]G01
| |--D. cochleata (Quenstedt 1858)GK02 (see below for synonymy)
| `--‘Rostellaria’ trifida Eudes-Deslongchamps 1842G01
`--Pugnellinae [Pugnellidae]BR17
| i. s.: Lispodesthes White 1877KB99, W77
| |--L. lingulifera White 1877W77
| |--L. nuptialis (White 1874) [=Anchura nuptialis]W77
| |--L. panda Stephenson 1953KB99
| `--L. patula Stephenson 1953KB99
| Tundora Stephenson 1941KB99, BR17 [Tundorinae]
| `--*T. tuberculata Stephenson 1941BR17
| TessarolaxKB99
| CeratosiphonKB99
|--Torgnellus peruvianus Ollson 1944KB99
|--Bizarrus Kiel & Bandel 1999KB99
| |--*B. dietrichi (Riedel 1932) [=Pugnellus dietrichi, P. dietrichei]KB99
| |--B. abnormalis (Wade 1926) [=Pugnellus abnormalis]KB99
| |--B. incertus (Riedel 1932) [=Pugnellus incertus]KB99
| `--‘Pugnellus’ nitidus Delpey 1948KB99
|--Pyktes Popenoe 1983KB99
| |--*P. aspris Popenoe 1983KB99
| |--P. bilobatus (Brebion 1956) [=Pugnellus bilobatus]KB99
| |--P. daiphron Popenoe 1983KB99
| |--P. hamulus Popenoe 1983KB99
| `--P. popenoei Kiel & Bandel 1999KB99
|--+--Tephlon Popenoe 1983KB99
| | `--*T. tumidus (Gabb 1860) [=Pugnellus tumidus]KB99
| `--+--AntarctodarwinellaKB99
| `--Conchothyra Hutton 1877KB99, SH12
| |--C. australis (Marshall 1916)SH12
| |--C. marshalli Trechmann 1917SH12
| `--C. parasitica Hutton 1877KB99
|--Gymnarus Gabb 1868KB99
| |--*G. manubriatus (Gabb 1864) [=Pugnellus manubriatus]KB99
| |--G. auriculatus (Woods 1906) [=Pugnellus auriculatus]KB99
| |--G. calcaris (Wade 1926)KB99
| |--G. congolensis (Brebion 1956) [=Pugnellus densatus var. congolensis]KB99
| |--G. contortus (Stoliczka 1868)KB99
| |--G. fraasi (Riedel 1932) [=Pugnellus fraasi]KB99
| |--G. granuliferus (Stoliczka 1868)KB99
| |--G. hauthali Wilckens 1910KB99
| `--G. yabei Nagao 1939KB99
`--+--Perustrombus Ollson 1944KB99
| |--*P. wheeleri Ollson 1944KB99
| |--P. africanus Kiel & Bandel 1999KB99
| |--‘Strombus’ dallianus White 1887KB99
| `--P. indicus Kiel & Bandel 1999KB99
`--Pugnellus Conrad 1858KB99
|--*P. densatus (Conrad 1858) [=Strombus densatus; incl. P. typicus Gabb 1877]BR17
|--P. bartheli Kiel & Bandel 1999KB99
|--P. goldmaniKB99
|--P. hauthali Wilckens 1905KB99
|--P. klitzschi Kiel & Bandel 1999KB99
`--P. uncatus Stoliczka 1868KB99

Aporrhaidae incertae sedis:
Mesochilotoma Seeley 1861P66
`--*M. striata Seeley 1861P66
Bicorempterus Gründel 2001G01
`--*B. elegans (Hudleston 1884) (see below for synonymy)G01
Pietteia Cossmann 1904G01
|--*P. hamus (Eudes-Deslongchamps 1842) [=Rostellaria hamus, Alaria hamus]G01
|--P. anglica Cox & Arkell 1950G01
|--P. callamus Gründel 2001G01
|--P. hudlestoni (Wilson 1887) [=Anchura (Pietteia) hudlestoni]TTE93
`--P. russiensisG01
Cuphotifer Piette 1876G01
Ueckeritzella Gründel 1998G01
`--U. mothsi Gründel 1998G01
Canepina Conti & Fischer 1981G01

Aporrhais serresianus (Michaud 1828) [=Rostellaria serresiana, Chenopus serresiana; incl. Aporrhais mcandreae, A. pescarbonis]N79

*Bicorempterus elegans (Hudleston 1884) [=Alaria bispinosa var. elegans, Dicroloma elegans; incl. A. bicornis Blake 1905-1907]G01

Dicroloma cochleata (Quenstedt 1858)GK02 [=Rostellaria cochleataG01, Alaria cochleataG01, Anchura (Dicroloma) cochleataG01, Dicroloma cochleatumG01]

Spiniloma Gründel, Nutzel & Schulbert 2009BR17 [=Spinigera d’Orbigny 1850 non Lesson 1842BR17; Spinigeridae, Spinigerinae, SpinilomatinaeBR05]

*Type species of generic name indicated


[A27] Allan, R. S. 1927. The geology and palaeontology of the Lower Waihao Basin, south Canterbury, New Zealand. Transactions and Proceedings of the New Zealand Institute 57: 265–309.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[G01] Gründel, J. 2001. Neritimorpha und weitere Caenogastropoda (Gastropoda) aus dem Dogger Norddeutschlands und des nordwestlichen Polens. Berliner Geowissenschaftliche Abhandlungen, Reihe E 36: 45–99.

[GK02] Gründel, J., & T. Kowalke. 2002. Palaeorissoinidae, a new family of marine and brackish water Rissooidea (Gastropoda, Littorinimorpha). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 226 (1): 43–57.

[KB99] Kiel, S., & K. Bandel. 1999. The Pugnellidae, a new stromboidean family (Gastropoda) from the Upper Cretaceous. Paläontologische Zeitschrift 73 (1–2): 47–58.

[N79] Norman, A. M. 1879. The Mollusca of the fiords near Bergen, Norway. Journal of Conchology 2: 8–77.

[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: an evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

[S98] Stilwell, J. D. 1998. Late Cretaceous Mollusca from the Chatham Islands, New Zealand. Alcheringa 22 (1): 29–85.

[SH12] Stilwell, J. D., & E. Håkansson. 2012. Survival, but…! New tales of ‘dead clade walking’ from Austral and Boreal post-K–T assemblages. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 795–810. Springer.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[W77] White, C. A. 1877. Report upon the invertebrate fossils collected in portions of Nevada, Utah, Colorado, New Mexico, and Arizona, by parties of the expeditions of 1871, 1872, 1873, and 1874. U.S. Geographical Surveys West of the One Hundredth Meridian 4 (1): 1–219, pls 1–21.

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