Javan hawk-eagle Nisaetus bartelsi, copyright Eko Prastyo.

Belongs within: Accipitridae.
Contains: Aquila.

The Aquilinae, booted eagles, are a cosmopolitan group of large eagles characterised by feathered tarsi (Lerner & Mindell 2005).

    |--+--Stephanoaetus coronatusJT12
    |  `--NisaetusJT12
    |       |  i. s.: N. florisJT12
    |       |--+--N. lanceolatusJT12 [=Spizaetus lanceolatusLM05]
    |       |  `--+--N. cirrhatusJT12 [=Spizaetus cirrhatusLM05]
    |       |     |    |--N. c. cirrhatusLM05
    |       |     |    `--'Spizaetus’ c. lineatusLM05
    |       |     `--N. philippensisJT12
    |       `--+--N. nanusJT12 [=Spizaetus nanusLM05]
    |          `--+--N. nipalensisJT12 [=Spizaetus nipalensisLM05]
    |             |    |--N. n. nipalensisI92
    |             |    `--'Spizaetus’ n. orientalis Temminck & Schlegel 1844I92
    |             `--+--N. albonigerJT12
    |                `--N. bartelsiJT12
    `--+--+--‘Spizaetus’ tyrannusJT12
       |  |    |--S. t. tyrannusE52
       |  |    `--S. t. serusE52
       |  `--+--Spizastur melanoleucusLM05 [=Spizaetus melanoleucusJT12]
       |     `--+--Oroaetus isidoriLM05 [=Spizaetus isidoriJT12]
       |        `--Spizaetus Vieillot 1816JT12, B94 [Spizaetinae]
       |             |--S. bellicosusFP64
       |             |--S. coronatusWH02
       |             |--S. grinnelliWH02
       |             `--S. ornatusJT12
       |                  |--S. o. ornatusE52
       |                  `--S. o. vicariusE52
       `--+--Lophotriorchis kieneriiJT12 [=Hieraaetus kieneriiLM05]
          `--+--Polemaetus Heine 1890JT12, M02
             |    `--P. bellicosusJT12
             `--+--+--Lophaetus occipitalisJT12
                |  `--+--Ictinaetus malayensisJT12
                |     `--+--‘Aquila’ hastataJT12 [=A. pomarina hastataLM05]
                |        `--+--‘Aquila’ clangaJT12
                |           `--‘Aquila’ pomarinaJT12
                   `--Hieraaetus Kaup 1844JT12, CC10 [incl. Harpagornis Haast 1872CC10]
                        |  i. s.: H. edwardsi (Sharpe 1899) (see below for synonymy)M02
                        |--H. wahlbergiLM05 [=Aquila wahlbergiJT12, Spizaetus wahlbergiLM05]
                        `--+--H. ayresiiJT12
                           `--+--H. morphnoides (Gould 1840)LM05, WS48 (see below for synonymy)
                              `--+--*H. pennatus (Gmelin 1788)CC10, JT12, CC10 (see below for synonymy)
                                 |--H. moorei (Haast 1872)CC10 (see below for synonymy)
                                 `--H. weiskeiJT12 [=H. morphnoides weiskeiLM05]

Hieraaetus edwardsi (Sharpe 1899) [=Eutolmaetus edwardsi, Aquila edwardsi, Nisaetus edwardsi, Aquila minuta Milne-Edwards 1871 non Brehm 1831]M02

Hieraaetus moorei (Haast 1872)CC10 [=*Harpagornis mooreiCC10, BKB15, Aquila mooreiCC10; incl. Ha. assimilis Haast 1874CC10]

Hieraaetus morphnoides (Gould 1840)LM05, WS48 [=Aquila morphnoidesWS48; incl. A. morphnoides coongani Mathews 1912WS48]

*Hieraaetus pennatus (Gmelin 1788)CC10, JT12, CC10 [=Falco pennatusCC10, Aquila pennataCC10; incl. A. minuta Brehm 1831M02]

*Type species of generic name indicated


[B94] Bock, W. J. 1994. History and nomenclature of avian family-group names. Bulletin of the American Museum of Natural History 222: 1–281.

[BKB15] Burleigh, J. G., R. T. Kimball & E. L. Braun. 2015. Building the avian tree of life using a large-scale, sparse supermatrix. Molecular Phylogenetics and Evolution 84: 53–63.

[CC10] Checklist Committee (OSNZ). 2010. Checklist of the Birds of New Zealand, Norfolk and Macquarie Islands, and the Ross Dependency, Antarctica 4th ed. Ornithological Society of New Zealand and Te Papa Press: Wellington.

[E52] Eisenmann, E. 1952. Annotated list of birds of Barro Colorado Island, Panama Canal Zone. Smithsonian Miscellaneous Collections 117 (5): 1–62.

[FP64] Fisher, J., & R. T. Peterson. 1964. The World of Birds: A comprehensive guide to general ornithology. Macdonald: London.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[JT12] Jetz, W., G. H. Thomas, J. B. Joy, K. Hartmann & A. Ø. Mooers. 2012. The global diversity of birds in space and time. Nature 491: 444–448.

[LM05] Lerner, H. R. L., & D. P. Mindell. 2005. Phylogeny of eagles, Old World vultures, and other Accipitridae based on nuclear and mitochondrial DNA. Molecular Phylogenetics and Evolution 37: 327–346.

[M02] Mlíkovský, J. 2002. Cenozoic Birds of the World. Part 1: Europe. Ninox Press: Praha.

[WS48] Whittell, H. M., & D. L. Serventy. 1948. A systematic list of the birds of Western Australia. Public Library, Museum and Art Gallery of Western Australia, Special Publication 1: 1–126.

[WH02] Worthy, T. H., & R. N. Holdaway. 2002. The Lost World of the Moa: Prehistoric life of New Zealand. Indiana University Press: Bloomington (Indiana).

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