Cybaeus nipponicus, from here.

Belongs within: Neocribellatae.

The Argyronetidae are a Holarctic group of spiders including the water spider Argyroneta aquatica and related terrestrial species. Argyroneta aquatica is the most truly aquatic spider, constructing an underwater ‘diving bell’ of silk in which it traps a bubble of air that allows it to spend almost its entire life submerged, making only occasional visits to the surface to collect fresh air. Terrestrial species have historically been recognised as a distinct family Cybaeidae but differences between ‘cybaeids’ and Argyroneta are largely related to the latter’s aquatic habits (Selden 2002).

Characters (from Selden 2002): Legs with unnotched trochanters, three tarsal claws, single row of tarsal trichobothria; ecribellate; spinnerets not in a transverse row, anterior spinnerets closely spaced, posterior spinnerets narrowed with very short distal segment.

<==Argyronetidae [Cybaeidae]
    |  i. s.: CybaeotaS02
    |         CybaeozygaS02
    |         Cybaeina minuta (Banks 1906)WZ08
    |--Argyronetinae [Argyroneteae]S02
    |    |  i. s.: Urquhartia Bryant 1933S02
    |    |--Vectaraneus yulei Selden 2001S02
    |    `--+--Elvina Thorell 1870S02
    |       |    `--*E. antiqua (von Heyden 1859) [=Argyroneta antiqua]S02
    |       `--Argyroneta Latreille 1804S02
    |            |--*A. aquatica (Clerck 1757)S02 [=Araneus aquaticusS02, Aranea aquaticaL58]
    |            |--A. bicolor Risso 1826R26
    |            |--A. longipes Heer 1865 [=Argyronecta (l. c.) longipes]S02
    |            |--A. palustris Risso 1826R26
    |            `--A. trilineataR26
    `--Cybaeus Koch 1868MG03 [Cybaeae, CybaeinaeS02]
         |--C. abchasicus Charitonov 1947MG03
         |--C. angustiarum Koch 1868MG03
         |--C. hesper Chamberlin & Ivie 1932S02
         |--C. hibaensisT07
         |--C. jilinensis (Song et al. 1993)WZ08
         |--C. nipponicusT07
         |--C. patritus Bishop & Crosby 1926S02
         |--C. reticulatus Simon 1886S06
         |--C. tetricusJD-S07
         `--C. tottoriensisT07
Nomina nuda: Cybaeus algidusS06
             Cybaeus borealisS06

*Type species of generic name indicated


[JD-S07] Jocqué, R., & A. S. Dippenaar-Schoeman. 2007. Spider Families of the World. Royal Museum for Central Africa: Tervuren (Belgium).

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[MG03] Marusik, Yu. M., & E. F. Guseinov. 2003. Spiders (Arachnida: Aranei) of Azerbaijan. 1. New family and genus records. Arthropoda Selecta 12 (1): 29–46.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

[S02] Selden, P. A. 2002. Missing links between Argyroneta and Cybaeidae revealed by fossil spiders. Journal of Arachnology 30 (2): 189–200.

[S06] Strand, E. 1906. Die arktischen Araneae, Opiliones und Chernetes. In: Römer, F., & F. Schaudinn (eds) Fauna Arctica. Eine Zusammenstellun der arktischen Tierformen, mit besonder Berücksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nördliche Eismeer im Jahre 1898 vol. 4 pp. 431–478. Gustav Fischer: Jena.

[T07] Tsurusaki, N. 2007. The Chugoku Mountains—a hotspot of geographical differentiation of species. Taxa 22: 3–14.

[WZ08] Wang, X.-P., & M.-S. Zhu. 2008. Himalmartensus, a new genus of the spider family Amaurobiidae from Nepal (Araneae). Journal of Arachnology 36 (2): 241–250.

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