Asilus crabroniformis, copyright António A. Gonçalves.

Belongs within: Asilidae.

The Asilinae are a group of robber flies with a long, bare stylus on the antenna. They are the most diverse subfamily of robber flies in temperate regions of the world but are less diverse in the tropics. Asilines are unusual among Diptera in the absence of  a crop or esophageal diverticulum (Fisher 2009). The larger number of species are assigned to the genus Asilus, including fossils going back to the Eocene (Wood 1981). Stout bristles are commonly present on the posterolateral margins of abdominal tergites 2 to 6 though these are absent in the New World genera Anarmostus and Wyliea (Fisher 2009).

Characters (from Fisher 2009): Antenna with apex of first flagellomere pointed, terminating in long, slender, arista-like stylus; apical flagellomere bare. Palpus one-segmented. Anatergite setose; postmetacoxal area membranous. Wing with R2+3 ending at R1, cell r1 closed; no appendicular vein on R4; crossvein m-cu absent; veins M3 and CuA1 joined together.

<==Asilinae [Asilini]F09
|--Promachella pilosaW81
|--Promachina trapezoidalisW81
|--Proctacanthella cacopilogaW81
|--Rhadiurgus variabilisW81
|--Polacantha (Echinitropis) xanthoceraF09
|--Antipalus varipesF09
|--Anarmostus iopterusF09
|--Prolatiforceps anonymaF09
|--Talamancasilus clavatusF09
|--Pamponerus germanicusJ61
|--Acanthopleura brunnipesJ61
|--Neoaratus herculesCM70
| |--M. frustraW81
| `--M. laphroidesE66
|--CerdistusCM91 [incl. NeomochtherusW81]
| |--C. elegansJ61
| `--‘Neomochtherus’ ochriventrisJ61
| |--P. albicepsF09
| `--P. arizonensisBT04
| |--W. chrysaugesF09
| `--W. mydasF09
| |--N. cyanurusV09
| `--N. orphneW81
| |--D. dasynotusJ61
| `--D. forcipulaK01
|--MachimusF09 [incl. TolmerusW81]
| |--M. achterbergi Tomasovic 2004F05
| |--M. gertschiF09
| |--M. paropusF09 [=Asilus (Machimus) paropusW81]
| |--M. perplexusJ61
| |--M. rusticusJ61
| `--M. soikai Janssens 1961J61
`--Asilus Linnaeus 1758L58
|--A. ater Linnaeus 1758L58
|--A. aureusL02
|--A. auriannulatusW81
|--A. barbarus Linnaeus 1758L58
|--A. bipartitus Macquart 1847E12
|--A. blantoniW81
|--A. crabroniformis Linnaeus 1758L58
|--A. cyanurusR13
|--A. flavopilosus Macquart 1847E12
|--A. forcipatus Linnaeus 1758L58
|--A. germanicus Linnaeus 1758L58
|--A. gibbosus Linnaeus 1758L58
|--A. gilvus Linnaeus 1758L58
|--A. hirtipes Macquart 1847E12
|--A. marginatus Linnaeus 1758L58
|--A. maurus Linnaeus 1758L58
|--A. morio Linnaeus 1758L58
|--A. oelandicus Linnaeus 1758L58
|--A. periscelis Macquart 1847E12
|--A. teutonusL02
|--A. tingitanus Boisduval 1835B35
`--A. tipuloides Linnaeus 1758L58

*Type species of generic name indicated


[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d’Urville. Faune entomologique de l’océan Pacifique, avec l’illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[BT04] Bybee, S. M., S. D. Taylor, C. R. Nelson & M. F. Whiting. 2004. A phylogeny of robber flies (Diptera: Asilidae) at the subfamilial level: molecular evidence. Molecular Phylogenetics and Evolution 30: 789–797.

[CM70] Colless, D. H., & D. K. McAlpine. 1970. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 656–740. Melbourne University Press.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[E66] Evans, H. E. 1966. The Comparative Ethology and Evolution of the Sand Wasps. Harvard University Press: Cambridge (Massachusetts).

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[F09] Fisher, E. M. 2009. Asilidae (robber flies, assassin flies, moscas cazadoras, moscas ladronas). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 585–632. NRC Research Press: Ottawa.

[J61] Janssens, E. 1961. Sur quelques asilides mediterranees. Boll. Mus. Civ. Venezia 14: 29–36.

[K01] Kertész, K. 1901. Legyek [Dipteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 179–201. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.

[R13] Reuter, O. M. 1913. Lebensgewohnheiten und Instinkte der Insekten bis zum Erwachen der sozialen Instinkte. R. Friedländer & Sohn: Berlin.

[V09] Verdcourt, B. (ed.) 2009. Additions to the wild fauna and flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.

[W81] Wood, G. C. 1981. Asilidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 549–573. Research Branch, Agriculture Canada.


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