Ctenitis hemsleyana, photographed by Robbin Moran.

Belongs within: Polypodiales.

The Aspidiaceae is a large but often poorly defined family of ferns. The separation or otherwise of this family from others such as Dryopteridaceae has often been contentious, but Wang (1965) pointed out the raised costa of the leaves in Aspidiaceae as a significant distinction.

Characters (from Wang 1965): Middle-sized, terrestrial ferns; rhizome erect or sub-erect, dictyostelic, scaly; leaves tufted, generally uniform, simple pinnate to tripinnatifid; veins free or anastomosing, forming areoles with or without free included veinlets; rachis and costa raised on the upper surface and generally covered with pale brown multicellular articulate hairs; sori round or in a few cases acrostichoid; indusia reniform, or in some genera absent; spores bilateral with perispore.

<==Aspidiaceae [Tectarioideae]
    |--Rhipidopteris peltataL54
    |--Polystichopsis Smith 1875L54, A61
    |    `--P. denticulataL54
    |--Aspidites Harris 1961 nec Peters 1876 (ICZN) nec Waagen 1895 (ICZN)BO02
    |    |--A. beckeriK-CB01
    |    |--A. delicatula Barale & Ouaja 2002BO02
    |    |--A. sewardiBO02
    |    `--A. thomasii Harris 1961C93
    |    |--T. coadunata (Smith) Christensen 1931B06 (see below for synonymy)
    |    |--T. hippocrepisJ87
    |    |--T. polymorpha (Wallich ex Hooker) Copel. 1907B06 (see below for synonymy)
    |    |--T. subcordataH03
    |    `--T. tabonensisH03
    |    |--C. caryotideum (Wallich) Presl 1836 (see below for synonymy)I88
    |    |--C. falcatum (L. f.) Presl 1836 [=Polypodium falcatum L. f. 1781]B06
    |    |--C. fortunei Smith 1866I88
    |    |--C. hookerianum (Presl) Christensen 1913 (see below for synonymy)I88
    |    `--C. macrophyllum (Makino) Tagawa 1934 [=Aspidium falcatum var. macrophyllum Makino 1902]I88
    `--Aspidium Swartz 1800M03
         |--A. cicutariumT-W89
         |--A. confluens [=Nephrodium confluens; incl. A. melanocaulon, Sagenia melanocaulon]B78
         |--A. cyathoidesBS-V28
         |--A. discolorBS-V28
         |--A. durvillaei Bory de Saint-Vincent 1828BS-V28
         |--A. falcatumT86
         |--A. filix-masN77
         |--A. fischeriS89
         |--A. fuscatumBS-V28
         |--A. lessonii Bory de Saint-Vincent 1828BS-V28
         |--A. leuzeanum [=Pleocnemia leuzeana]T-W89
         |--A. membranaceumT-W89
         |--A. meniscioidesBS-V28
         |--A. meyeriS89
         |--A. multifidumD03
         |--A. orbiculatumD03
         |--A. pachyphyllumT-W89
         |--A. serraB78
         |--A. singaporianumT-W89
         |--A. subtriphyllumT-W89
         |--A. trifoliatumBS-V28
         |--A. variolosumT-W89
         `--A. vastumT-W89

Nomen nudum: Aspidium apenninicum Squinabol 1889S89
             Aspidium oligocenicum Squinabol 1889S89
             Aspidium pareti Squinabol 1889S89

Cyrtomium caryotideum (Wallich) Presl 1836 [=Aspidium caryotideum Wallich ex Hooker & Greville 1828, C. falcatum var. caryotideum (Wallich) Bedd. 1883]I88

Cyrtomium hookerianum (Presl) Christensen 1913 [=Lastrea hookeriana Presl 1836; incl. Aspidium caducum (Presl) Wallich ex Clarke 1880]I88

Tectaria coadunata (Smith) Christensen 1931B06 [=Aspidium coadunatum Wallich ex Hooker & Greville 1831I88, Nephrodium cicutarium var. coadunatum (Wallich) Clarke 1880I88, Sagenia coadunata Smith in Hooker 1841B06]

Tectaria polymorpha (Wallich ex Hooker) Copel. 1907B06 [=Aspidium polymorphum Wallich 1828B06, Nephrodium polymorphum (Wallich) Hooker & Baker 1865I88]

*Type species of generic name indicated


[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).

[BO02] Barale, G., & M. Ouaja. 2002. La biodiversité végétale des gisements d’âge Jurassique supérieur-Crétacé inférieur de Merbah El Asfer (Sud-Tunisien). Cretaceous Research 23: 707–737.

[B78] Bentham, G. 1878. Flora Australiensis: A description of the plants of the Australian Territory vol. 7. Roxburghiaceae to Filices. L. Reeve & Co.: London.

[B06] Biswas, A. 2006. Pteridophytes of Indian Botanic Garden, Howrah. Bulletin of the Botanical Survey of India 48: 175–188.

[BS-V28] Bory de Saint-Vincent, J. B. 1828. Voyage Autour du Monde, Exécuté par Ordre du Roi, Sur la Corvette de Sa Majesté, La Coquille, pendant les années 1822, 1823, 1824 et 1825. Botanique. Cryptogamie. Arthus Bertrand: Paris.

[C93] Cleal, C. J. 1993. Pteridophyta. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 779–794. Chapman & Hall: London.

[D03] Dusén, P. 1903. The vegetation of western Patagonia. In: Scott, W. B. (ed.) Reports of the Princeton University Expeditions to Patagonia, 18961899 vol. 8. Botany pp. 1–34. The University: Princeton (New Jersey).

[H03] Heads, M. 2003. Ericaceae in Malesia: vicariance biogeography, terrane tectonics and ecology. Telopea 10 (1): 311–449.

[I88] Iwatsuki, K. 1988. An enumeration of the pteridophytes of Nepal. In: Ohba, H., & S. B. Malla (eds) The Himalayan Plants vol. 1. The University Museum, University of Tokyo, Bulletin 31: 231–339.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State MuseumBiological Sciences 32 (1): 1–136.

[K-CB01] Konijnenburg-van Cittert, H. van, & K. Bandel. 2001. Jurassic plants from Djebel Tih, Sinai. Mitteilungen aus dem Geologisch-Paläontologischen Institut der Universität Hamburg 85: 47–64.

[L54] Lötschert, W. 1954. Beitrag zur Pteridophyten-Flora von Mittel-Amerika. Senckenbergiana Biologica 35 (1-2): 109–119.

[M03] Macloskie, G. 1903. Pteridophyta, ferns and fernlike plants. In: Scott, W. B. (ed.) Reports of the Princeton University Expeditions to Patagonia, 18961899 vol. 8. Botany pp. 127–138. The University: Princeton (New Jersey).

[N77] Niedbała, W. 1977. Badania biocenotyczne nad mechowcami (Acari, Oribatei) Masywu Ślęży. Badania Fizjograficzne nad Polską Zachodnią, Seria C—Zoologia 30: 47–73.

[S89] Squinabol, S. 1889. Res Ligusticae. VII.—Cenno preliminare sulla flora fossile di Santa Giustina. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 73–76.

[T-W89] Tenison-Woods, J. E. 1889. On the vegetation of Malaysia. Proceedings of the Linnean Society of New South Wales, series 2, 4 (1): 9–106, pls 1–9.

[T86] Trebeck, P. N. 1886. Mount Wilson and its ferns. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 491–496.

Wang C.-H. 1965. A taxonomical study of the family Aspidiaceae from the mainland of Asia. Acta Phytotaxonomica Sinica 10 (2): 121–130.

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