Lyon’s pentachaeta Pentachaeta lyonii, from the Pacific Southwest Region USFWS.

Belongs within: Asteroideae.
Contains: Aster, Olearia, Calotis, Erigeron, Vittadinia, Hazardia, Bellidinae, Celmisia, Baccharis, Grindelia, Chrysothamnus, Solidago, Ericameria.

The composite-flowered tribe Astereae includes asters (Aster), goldenrods (Solidago) and related taxa. Members of the group are distinguished by the morphology of the fertile disc-flowers, in which the marginal stigmatic lines do not reach the summits of the style-branches. Within the Astereae, members of the subtribe Asterinae have a well-developed pappus, as opposed to the reduced pappus of the Bellidinae (Black & Robertson 1965). Pleurophyllum species are robust perennial herbs found on the Subantarctic islands of New Zealand (Allan 1961). Conyza, the fleabanes, are erect herbs or shrubs growing up to two metres in height, some species of which are notable as invasive weeds. Minuria is an Australian genus of mostly small undershrubs with flower-heads borne on short peduncles; the receptacle is naked and ray-flowers are borne in two or more rows (Black & Robertson 1965).

Characters (from Black & Robertson 1965): Style-branches of fertile disc-flowers flattish, with two marginal papillose stigmatic lines along lower part of branches and long or short non-stigmatic appendage at summit bearing short collecting hairs on outside; flower-heads heterogamous radiate; receptacle usually naked; pappus consisting of bristles, awns or scales, rarely absent; leaves alternate.

    |--Doellingeria umbellataRU03
    |--+--Gundlachia corymbosaRU03
    |  `--XylothamiaRU03
    |       |--X. parrasana [=Haplopappus parrasanus]RU03
    |       |--‘Haplopappus’ sonoriensisRU03
    |       |--X. trianthaRU03
    |       `--X. truncataRU03
    `--+--+--‘Chrysothamnus’ gramineusRU03 [incl. Petradoria discoideaH93]
       |  `--+--Sericocarpus tortifoliusRU03
       |     `--+--ChrysothamnusRU03
       |        `--+--Chrysoma pauciflosculosaRU03
       |           `--SolidagoRU03
          `--+--+--Rigiopappus leptocladusRU03
             |  `--Tracyina rostrataRU03
                  |--P. alsinoides [=Chaetopappa alsinoides]H93
                  |--P. aurea [=Chaetopappa aurea]H93
                  |--P. bellidiflora [=Chaetopappa bellidiflora]H93
                  |--P. exilis [=Chaetopappa exilis]H93
                  |    |--P. e. ssp. exilisH93
                  |    `--P. e. ssp. aeolicaH93
                  |--P. fragilis [=Chaetopappa fragilis]H93
                  `--P. lyonii [=Chaetopappa lyonii]H93
Astereae incertae sedis:
    |--Achnophora tateiBR65
    |    |--P. cuneifoliaBR65
    |    `--P. nanaBR65
         |--M. annua [=Minuriella annua]BR65
         |--M. cunninghamiiBR65
         |--M. denticulataBR65
         |--M. integerrimaBR65
         |--M. leptophyllaBR65
         |--M. macrorhizaLK14
         |--M. rigidaBR65
         `--M. suaedifoliaBR65
  Stenactis annua [=Aster annuus]C55
    |--H. coronopifoliusD03
    |--H. macleaniiM69a
    |--H. odontolepis Moran 1969M69a
    |--H. resinosusPS96
    |--H. rosaricus Moran 1969M69a
    |--H. rubiginosus [=Sideranthus rubiginosus]G05
    |--H. spinulosusA72
    |--H. tenuisectusLW70
    `--H. wigginsiiA72
  Pleurophyllum Hooker 1844A61
    |--P. criniferum Hooker 1844A61 (see below for synonymy)
    |--P. hookeri Buchan. 1884A61 [incl. P. gilliesianum Kirk 1891C06, P. hookerianum Kirk 1891C06]
    `--P. speciosum Hooker 1844A61 [incl. Aster pleurophyllum Muell. 1865ME70]
  Pachystegia Cheeseman 1925A61
    `--P. insignis (Hooker) Cheeseman 1925 (see below for synonymy)A61
         |--P. i. var. insignisA61
         `--P. i. var. minor Cheeseman 1925 [=Olearia insignis var. minor Cheeseman 1916]A61
    |--G. californica [incl. G. bracteata]H93
    |--G. lacidaBT72
    |--G. microcephalaH93
    `--G. sarothraeCL86
  Chiliotrichium diffusumS06, CL85
    |--C. albidaGK00
    |--C. apurensisJ87
    |--C. bilboanaH93
    |--C. bonariensisY98
    |--C. canadensisAGF98
    |    |--C. c. var. canadensisJ87
    |    `--C. c. var. pusillaJ87
    |--C. coulteriH93
    `--C. floribundaH93
  Chrysocoma puberulaS06, CV06
  Heterothalamus nivalisS06, D03

Pachystegia insignis (Hooker) Cheeseman 1925 [=Olearia insignis Hooker 1855; incl. O. marginata Colenso 1883]A61

Pleurophyllum criniferum Hooker 1844A61 [=Aster criniferum (Hooker) Muell. 1865ME70; incl. P. hombronii Decne 1853A61, Albinia oresigenesa Homb. & Jacq. ex Decne 1853A61, P. oresigenesumC06]

*Type species of generic name indicated


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[BR65] Black, J. M., & E. L. Robertson. 1965. Flora of South Australia. Part IV. Oleaceae–Compositae. W. L. Hawes, Government Printer: Adelaide.

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[C06] Cheeseman, T. F. 1906. Manual of the New Zealand Flora. John Mackay, Government Printer: Wellington.

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[CL86] Collins, E. I., & R. W. Lichvar. 1986. Vegetation inventory of current and historic black-footed ferret habitat in Wyoming. Great Basin Naturalist Memoirs 8: 85–93.

[CV06] Craven, P., & P. Vorster. 2006. Patterns of plant diversity and endemism in Namibia. Bothalia 36 (2): 175–189.

[D03] Dusén, P. 1903. The vegetation of western Patagonia. In: Scott, W. B. (ed.) Reports of the Princeton University Expeditions to Patagonia, 1896–1899 vol. 8. Botany pp. 1–34. The University: Princeton (New Jersey).

[GK00] Gibson, N., & G. J. Keighery. 2000. Flora and vegetation of the Byenup-Muir reserve system, south-west Western Australia. CALMScience 3 (3): 323–402.

[G05] Grissell, E. E. 2005. A review of North American species of Microdontomerus Crawford (Torymidae: Hymenoptera). Journal of Hymenoptera Research 14 (1): 22–65.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.

[LW70] Lowe, C. H., J. W. Wright, C. J. Cole & R. L. Bezy. 1970. Natural hybridization between the teiid lizards Cnemidophorus sonorae (parthenogenetic) and Cnemidophorus tigris (bisexual). Systematic Zoology 19 (2): 114–127.

[LK14] Lyons, M. N., G. J. Keighery, L. A. Gibson & T. Handasyde. 2014. Flora and vegetation communities of selected islands off the Kimberley coast of Western Australia. Records of the Western Australian Museum Supplement 81: 205–244.

[ME70] Moore, L. B., & E. Edgar. 1970. Flora of New Zealand vol. 2. Indigenous Tracheophyta: Monocotyledones except Gramineae. A. R. Shearer, Government Printer: Wellington (New Zealand).

[M-FR08] Morais-Filho, J. C., & G. Q. Romero. 2008. Microhabitat use by Peucetia flava (Oxyopidae) on the glandular plant Rhyncanthera dichotoma (Melastomataceae). Journal of Arachnology 36 (2): 374–378.

[M69] Moran, R. 1969. Five new taxa of Haplopappus (Compositae) from Baja California, Mexico. Transactions of the San Diego Society of Natural History 15 (11): 149–164.

[N10] Norrbom, A. L. 2010. Tephritidae (fruit flies, moscas de frutas). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 909–954. NRC Research Press: Ottawa.

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[RU03] Roberts, R. P., & L. E. Urbatsch. 2003. Molecular phylogeny of Ericameria (Asteraceae, Astereae) based on nuclear ribosomal 3´ ETS and ITS sequence data. Taxon 52: 209–228.

[S06] Stuckert, T. 1906. Distribución geográfica de la flora Argentina. Géneros de la familia de las compuestas. Anales del Museo Nacional de Buenos Aires, serie 3, 6: 303–309.

[Y98] Yannitsaros, A. 1998. Additions to the flora of Kithira (Greece) I. Willdenowia 28: 77–94.

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