Astereae

 Lyon’s pentachaeta Pentachaeta lyonii, from the Pacific Southwest Region USFWS.

Belongs within: Asteroideae.
Contains: Aster, Olearia, Calotis, Erigeron, Vittadinia, Hazardia, Bellidinae, Celmisia, Baccharis, Grindelia, Chrysothamnus, Solidago, Ericameria.

The composite-flowered tribe Astereae includes asters (Aster), goldenrods (Solidago) and related taxa. Members of the group are distinguished by the morphology of the fertile disc-flowers, in which the marginal stigmatic lines do not reach the summits of the style-branches. Within the Astereae, members of the subtribe Asterinae have a well-developed pappus, as opposed to the reduced pappus of the Bellidinae (Black & Robertson 1965). Pleurophyllum species are robust perennial herbs found on the Subantarctic islands of New Zealand (Allan 1961). Conyza, the fleabanes, are erect herbs or shrubs growing up to two metres in height, some species of which are notable as invasive weeds. Minuria is an Australian genus of mostly small undershrubs with flower-heads borne on short peduncles; the receptacle is naked and ray-flowers are borne in two or more rows (Black & Robertson 1965).

Characters (from Black & Robertson 1965): Style-branches of fertile disc-flowers flattish, with two marginal papillose stigmatic lines along lower part of branches and long or short non-stigmatic appendage at summit bearing short collecting hairs on outside; flower-heads heterogamous radiate; receptacle usually naked; pappus consisting of bristles, awns or scales, rarely absent; leaves alternate.

<==Astereae
    |--Doellingeria umbellataRU03
    |--+--Gundlachia corymbosaRU03
    |  `--XylothamiaRU03
    |       |--X. parrasana [=Haplopappus parrasanus]RU03
    |       |--‘Haplopappus’ sonoriensisRU03
    |       |--X. trianthaRU03
    |       `--X. truncataRU03
    `--+--+--‘Chrysothamnus’ gramineusRU03 [incl. Petradoria discoideaH93]
       |  `--+--Sericocarpus tortifoliusRU03
       |     `--+--ChrysothamnusRU03
       |        `--+--Chrysoma pauciflosculosaRU03
       |           `--SolidagoRU03
       `--+--EricameriaRU03
          `--+--+--Rigiopappus leptocladusRU03
             |  `--Tracyina rostrataRU03
             `--PentachaetaRU03
                  |--P. alsinoides [=Chaetopappa alsinoides]H93
                  |--P. aurea [=Chaetopappa aurea]H93
                  |--P. bellidiflora [=Chaetopappa bellidiflora]H93
                  |--P. exilis [=Chaetopappa exilis]H93
                  |    |--P. e. ssp. exilisH93
                  |    `--P. e. ssp. aeolicaH93
                  |--P. fragilis [=Chaetopappa fragilis]H93
                  `--P. lyonii [=Chaetopappa lyonii]H93
Astereae incertae sedis:
  AsterinaeBR65
    |--AsterBR65
    |--Achnophora tateiBR65
    |--OleariaBR65
    |--CalotisBR65
    |--ErigeronBR65
    |--VittadiniaBR65
    |--PodocomaBR65
    |    |--P. cuneifoliaBR65
    |    `--P. nanaBR65
    `--MinuriaBR65
         |--M. annua [=Minuriella annua]BR65
         |--M. cunninghamiiBR65
         |--M. denticulataBR65
         |--M. integerrimaBR65
         |--M. leptophyllaBR65
         |--M. macrorhizaLK14
         |--M. rigidaBR65
         `--M. suaedifoliaBR65
  HazardiaRU03
  Stenactis annua [=Aster annuus]C55
  HaplopappusM-FR08
    |--H. coronopifoliusD03
    |--H. macleaniiM69a
    |--H. odontolepis Moran 1969M69a
    |--H. resinosusPS96
    |--H. rosaricus Moran 1969M69a
    |--H. rubiginosus [=Sideranthus rubiginosus]G05
    |--H. spinulosusA72
    |--H. tenuisectusLW70
    `--H. wigginsiiA72
  BellidinaeBR65
  Pleurophyllum Hooker 1844A61
    |--P. criniferum Hooker 1844A61 (see below for synonymy)
    |--P. hookeri Buchan. 1884A61 [incl. P. gilliesianum Kirk 1891C06, P. hookerianum Kirk 1891C06]
    `--P. speciosum Hooker 1844A61 [incl. Aster pleurophyllum Muell. 1865ME70]
  CelmisiaA61
  Pachystegia Cheeseman 1925A61
    `--P. insignis (Hooker) Cheeseman 1925 (see below for synonymy)A61
         |--P. i. var. insignisA61
         `--P. i. var. minor Cheeseman 1925 [=Olearia insignis var. minor Cheeseman 1916]A61
  BaccharisN10
  GrindeliaS06
  GutierreziaS06
    |--G. californica [incl. G. bracteata]H93
    |--G. lacidaBT72
    |--G. microcephalaH93
    `--G. sarothraeCL86
  HysterionicaS06
  NardophyllumS06
  ChiliophyllumS06
  EgletesS06
  LeucopsidiumS06
  SommerfeldtiaS06
  Chiliotrichium diffusumS06, CL85
  ConyzaS06
    |--C. albidaGK00
    |--C. apurensisJ87
    |--C. bilboanaH93
    |--C. bonariensisY98
    |--C. canadensisAGF98
    |    |--C. c. var. canadensisJ87
    |    `--C. c. var. pusillaJ87
    |--C. coulteriH93
    `--C. floribundaH93
  Chrysocoma puberulaS06, CV06
  Heterothalamus nivalisS06, D03

Pachystegia insignis (Hooker) Cheeseman 1925 [=Olearia insignis Hooker 1855; incl. O. marginata Colenso 1883]A61

Pleurophyllum criniferum Hooker 1844A61 [=Aster criniferum (Hooker) Muell. 1865ME70; incl. P. hombronii Decne 1853A61, Albinia oresigenesa Homb. & Jacq. ex Decne 1853A61, P. oresigenesumC06]

*Type species of generic name indicated

References

[AGF98] Abd El-Ghani, M. M., & A. G. Fahmy. 1998. Composition of and changes in the spontaneous flora of Feiran Oasis, S Sinai, Egypt, in the last 60 years. Willdenowia 28: 123–134.

[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).

[A72] Atchley, W. R. 1972. The chromosome karyotype in estimation of lineage relationships. Systematic Zoology 21 (2): 199–209.

[BT72] Baker, E. W., & D. M. Tuttle. 1972. New species and further notes on the Tetranychoidea mostly from the southwestern United States (Acarina: Tetranychidae and Tenuipalpidae). Smithsonian Contributions to Zoology 116: 1–37.

[BR65] Black, J. M., & E. L. Robertson. 1965. Flora of South Australia. Part IV. Oleaceae–Compositae. W. L. Hawes, Government Printer: Adelaide.

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[C06] Cheeseman, T. F. 1906. Manual of the New Zealand Flora. John Mackay, Government Printer: Wellington.

[CL85] Cokendolpher, J. C., & D. Lanfranco L. 1985. Opiliones from the Cape Horn Archipelago: new southern records for harvestmen. Journal of Arachnology 13: 311–319.

[CL86] Collins, E. I., & R. W. Lichvar. 1986. Vegetation inventory of current and historic black-footed ferret habitat in Wyoming. Great Basin Naturalist Memoirs 8: 85–93.

[CV06] Craven, P., & P. Vorster. 2006. Patterns of plant diversity and endemism in Namibia. Bothalia 36 (2): 175–189.

[D03] Dusén, P. 1903. The vegetation of western Patagonia. In: Scott, W. B. (ed.) Reports of the Princeton University Expeditions to Patagonia, 1896–1899 vol. 8. Botany pp. 1–34. The University: Princeton (New Jersey).

[GK00] Gibson, N., & G. J. Keighery. 2000. Flora and vegetation of the Byenup-Muir reserve system, south-west Western Australia. CALMScience 3 (3): 323–402.

[G05] Grissell, E. E. 2005. A review of North American species of Microdontomerus Crawford (Torymidae: Hymenoptera). Journal of Hymenoptera Research 14 (1): 22–65.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.

[LW70] Lowe, C. H., J. W. Wright, C. J. Cole & R. L. Bezy. 1970. Natural hybridization between the teiid lizards Cnemidophorus sonorae (parthenogenetic) and Cnemidophorus tigris (bisexual). Systematic Zoology 19 (2): 114–127.

[LK14] Lyons, M. N., G. J. Keighery, L. A. Gibson & T. Handasyde. 2014. Flora and vegetation communities of selected islands off the Kimberley coast of Western Australia. Records of the Western Australian Museum Supplement 81: 205–244.

[ME70] Moore, L. B., & E. Edgar. 1970. Flora of New Zealand vol. 2. Indigenous Tracheophyta: Monocotyledones except Gramineae. A. R. Shearer, Government Printer: Wellington (New Zealand).

[M-FR08] Morais-Filho, J. C., & G. Q. Romero. 2008. Microhabitat use by Peucetia flava (Oxyopidae) on the glandular plant Rhyncanthera dichotoma (Melastomataceae). Journal of Arachnology 36 (2): 374–378.

[M69] Moran, R. 1969. Five new taxa of Haplopappus (Compositae) from Baja California, Mexico. Transactions of the San Diego Society of Natural History 15 (11): 149–164.

[N10] Norrbom, A. L. 2010. Tephritidae (fruit flies, moscas de frutas). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 909–954. NRC Research Press: Ottawa.

[PS96] Pike, K. S., P. Starý, R. Miller, D. Allison, L. Boydston, G. Graf & T. Miller. 1996. New species and host records of aphid parasitoids (Hymenoptera: Braconidae: Aphidiinae) from the Pacific northwest, U.S.A. Proceedings of the Entomological Society of Washington 98 (3): 570–591.

[RU03] Roberts, R. P., & L. E. Urbatsch. 2003. Molecular phylogeny of Ericameria (Asteraceae, Astereae) based on nuclear ribosomal 3´ ETS and ITS sequence data. Taxon 52: 209–228.

[S06] Stuckert, T. 1906. Distribución geográfica de la flora Argentina. Géneros de la familia de las compuestas. Anales del Museo Nacional de Buenos Aires, serie 3, 6: 303–309.

[Y98] Yannitsaros, A. 1998. Additions to the flora of Kithira (Greece) I. Willdenowia 28: 77–94.

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