Atrypida

 Anoplotheca venusta, from here.

Belongs within: Rhynchonellata.
Contains: Zygospiridae, Lissatrypidae, Atrypidae, Davidsoniidae.

The Atrypida are a group of spiralia-bearing brachiopods known from the Middle Ordovician to Upper Devonian. Members of this group may be divided between the Atrypacea, in which the spiralia are directed medially or dorsomedially, and the Dayiacea, in which the spiralia are directed ventrally or laterally, or are planispiral parallel to the median plane. Within the Dayiacea, members of the Dayiidae have smooth shells whereas members of the other families are variously ornamented (Boucot et al. 1965).

Characters (from Boucot et al. 1965, as Atrypidina): Impunctate, mostly biconvex spire-bearing brachiopods, commonly with narrow cardinal margin. Interarea low, obsolescent, or absent. Pedicle-valve beak may be truncated by foramen or foramen may reside within delthyrium. Spiralia directed medially, dorsomedially, or laterally; crura projecting to join primary lamellae of spiralia at posteromedial position, or deflected laterally more or less parallel to plane of commissure to join primary lamellae in posterolateral position; simple jugum commonly present connecting primary lamellae.

<==Atrypida
    |--Zygospiroidea [Zygospiridina]HB93
    |    |--ZygospiridaeHB93
    |    `--Tuvaella Chernyshev 1937WW65 [TuvaellidaeHB93]
    |         |--*T. rackovskii Chernyshev 1937WW65
    |         `--T. giganteaHB93
    |              |--T. g. giganteaHB93
    |              `--T. g. buchtarmaensis Kulkov & Kozlov 1978HB93
    |--Lissatrypoidea [Lissatrypidina]HB93
    |    |--LissatrypidaeBJ65
    |    `--ProtozygidaeHB93
    |         |--ManespiraHB93
    |         |--Idiospira Cooper 1956HB93, BJ65
    |         |    `--*I. panderi (Billings 1859) [=Camerella panderi]BJ65
    |         `--Protozyga Hall & Clarke 1893BJ65
    |              |--*P. exigua (Hall 1847) [=Atrypa exigua]BJ65
    |              `--P. elongataWR65
    |--AtrypidinaHB93
    |    |--Punctatrypa Havlíček 1953BJ65 [Punctatrypidae, Punctatrypinae, PunctatrypoideaHB93]
    |    |    |--*P. nalivkini Havlíček 1953BJ65
    |    |    |--P. granuliferaBJ65
    |    |    |--P. siehli Johnson 1975HB93
    |    |    `--P. tumidula (Khodalevich 1951)HB93
    |    `--Atrypoidea [Atrypacea]HB93
    |         |  i. s.: Loilemia Reed 1936BJ65
    |         |           `--*L. proxima Reed 1936BJ65
    |         |--AtrypidaeM03
    |         `--Atrypinidae [Atrypininae]HB93
    |              |--Sulcatospira parva (Rukavishnikova 1970)HB93
    |              `--Atrypina Hall & Clarke 1893 [Atrypinidae, Atrypininae]BJ65
    |                   |--*A. imbricata (Hall 1857) [=Leptocoelia imbricata]BJ65
    |                   |--A. hamiBJ65
    |                   `--A. prosimpsoni Johnson et al. 1973HB93
    `--DavidsoniideaHB93
         |--DavidsoniidaeM-WW65
         `--+--EospirigerinaB75
            |--Plectatrypa Schuchert & Cooper 1930B75, BJ65
            |    `--*P. ‘imbricata’ (Sowerby in Murchison 1839) [=Terebratula imbricata non Sowerby 1822]BJ65
            |--Spirigerina d’Orbigny 1849B75, BJ65
            |    |--*S. marginalis (Dalman 1828) [=Terebratula marginalis]BJ65
            |    |--S. concentricaH04
            |    `--S. supramarginalisH79
            |         |--S. s. supramarginalisTY12
            |         `--S. s. sibiricaTY12
            `--+--‘Plectatrypa’ henningsmoeni Boucot & Johnson 1967B75
               `--Karpinskiidae [Palaferelloidea]HB93
                    |  i. s.: Tectatrypa tectiformis (Chernyshev 1893)HB93
                    |--Eokarpinskia nalivkini (Nikiforova 1937)B75, HB93
                    `--+--Karpinskia Chernyshev 1885B75, BJ65 [incl. Notoconchidium Gill 1950BJ65]
                       |    |--*K. conjugula Chernyshev 1885BJ65
                       |    |--‘Pentamerus’ tasmaniensis Etheridge 1883 [=*Notoconchidium tasmaniense]BJ65
                       |    `--K. thomasi (Gill 1951) [=Notoconchidium thomasi]BJ65
                       `--+--Toquimaella kayiB75, N79
                          `--+--MimatrypaB75
                             |--Vagrania Alekseeva 1959B75, BJ65 [=Dentatrypa Breivel 1959BJ65]
                             |    |--*V. (Vagrania) kolymensis (Nalivkin 1936) (see below for synonymy)BJ65
                             |    `--V. (Desatrypa)HB93
                             |         |--V. (D.) desquamata (Sowerby 1840)HB93
                             |         `--V. (D.) globosa (Leidhold 1928)HB93
                             `--Palaferellidae [Palaferellinae]HB93
                                  |--Falsatrypa Havlíček 1956BJ65
                                  |    `--*F. admiranda Havlíček 1956BJ65
                                  |--Pseudogruenewaldtia Rzhonsnitskaya 1960NU83
                                  |    `--*P. tschernyschewi (Rzhonsnitskaya 1060)NU83
                                  `--Gruenewaldtia Chernyshev 1885B75, BJ65 [incl. Palaferella Spriestersbach 1942BJ65]
                                       |--*G. latilinguis (Schnur 1851) [=Terebratula latilinguis]BJ65
                                       |--G. gregeri [=Atrypa gregeri]NU83
                                       `--G. sibirica Ivanova 1956HB93

*Vagrania (Vagrania) kolymensis (Nalivkin 1936) [=Atrypa kolymensis, *Dentatrypa kolymensis]BJ65

*Type species of generic name indicated

References

[B75] Boucot, A. J. 1975. Evolution and Extinction Rate Controls. Elsevier Scientific Publishing Company.

[BJ65] Boucot, A. J., J. G. Johnson, C. W. Pitrat & R. D. Staton. 1965. Spiriferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt H. Brachiopoda vol. 2 pp. H632–H728. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas Press: Lawrence (Kansas).

[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.

[HB93] Harper, D. A. T., C. H. C. Brunton, L. R. M. Cocks, P. Copper, E. N. Doyle, A. L. Jeffrey, E. F. Owen, M. A. Parkes, L. E. Popov & C. D. Prosser. 1993. Brachiopoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 427–462. Chapman & Hall: London.

[H79] House, M. R. 1979. Devonian in the Eastern Hemisphere. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A183–A217. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[M03] Modzalevskaya, T. L. 2003. Silurian and Devonian brachiopods from Severnaya Zemlya (Russian Arctic). Geodiversitas 25 (1): 73–107.

[M-WW65] Muir-Wood, H., & A. Williams. 1965. Strophomenida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt H. Brachiopoda vol. 1 pp. H361–H521. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas Press: Lawrence (Kansas).

[N79] Norris, A. W. 1979. Devonian in the Western Hemisphere. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A218–A253. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[NU83] Norris, A. W., & T. T. Uyeno. 1983. Biostratigraphy and paleontology of middle-upper Devonian boundary beds, Gypsum Cliffs area, northeastern Alberta. Geological Survey of Canada Bulletin 313: 1–65.

[TY12] Talent, J. A., Y. A. Yokin, V. N, Yolkina, T. P. Kipriyanova & R. T. Gratsianova. 2012. Changes in the pattern of brachiopod biogeography in northern Asia through early and middle Devonian times. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 821–829. Springer.

[WR65] Williams, A., & A. J. Rowell. 1965. Morphology. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt H. Brachiopoda vol. 1 pp. H57–H138. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas Press: Lawrence (Kansas).

[WW65] Williams, A., & A. D. Wright. 1965. Orthida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt H. Brachiopoda vol. 1 pp. H299–H359. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas Press: Lawrence (Kansas).

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