Azhdarchoidea

Tupuxuara leonardii in the American Museum of Natural History, copyright Ole Kinderhook.

Belongs within: Pterodactyloidea.
Contains: Azhdarchidae.

The Azhdarchoidea are a Cretaceous lineage of pterosaurs, characterised by toothless jaws, very large nasoantorbital fenestrae extending above the eye sockets, short wing fingers with truncated distal wing phalanges, and long legs (Witton 2013).

<==AzhdarchoideaW13
    |  i. s.: Palaeornis cliftii Mantell 1844 (n. d.) (see below for synonymy)RK13
    |--Tapejaridae [Tapejarinae]W13
    |    |--Europejara olcadesorumW13
    |    |--TupandactylusW13
    |    |    |--T. imperatorW13
    |    |    `--T. navigans (Frey, Martill & Buchy 2003)W13, FMB03 [=Tapejara navigansW13]
    |    |--Tapejara Kellner 1989FMB03
    |    |    |--T. imperator Campos & Kellner 1997FMB03
    |    |    `--T. wellnhoferi Kellner 1989FMB03
    |    `--Sinopterus [incl. Huaxiapterus]W13
    |         |--‘Huaxiapterus’ benxiensisW13
    |         |--‘Huaxiapterus’ corollatusW13
    |         |--S. dongi [incl. S. gui]W13
    |         `--S. jii [=*Huaxiapterus jii]W13
    `--NeoazhdarchiaW13
         |  i. s.: LonchodectidaeW13
         |           |--Prejanopterus curvirostraW13
         |           |--Unwindia trigonus Martill 2011W13
         |           `--Lonchodectes Hooley 1914RK13
         |                `--*L. compressirostris (Owen 1851) (n. d.) (see below for synonymy)RK13
         |--ChaoyangopteridaeW13
         |    |--Chaoyangopterus zhangiW13
         |    |--Lacusovagus magnificensW13
         |    |--Eopteranodon liiW13
         |    |--Shenzhoupterus chaoyangensisW13
         |    |--Microtuban altivolansW13
         |    |--Jidapterus Dong et al. 2003W13, A08
         |    |    `--J. edentusW13
         |    `--Eoazhdarcho Lu & Ji 2005W13, A08
         |         `--E. liaoxiensisW13
         `--+--AzhdarchidaeW13
            `--ThalassodromidaeW13
                 |  i. s.: ‘Santanadactylus’ spixi Wellnhofer 1985W13, U03
                 |--Thalassodromeus Kellner & Campos 2002KC02
                 |    `--*T. sethi Kellner & Campos 2002KC02
                 `--Tupuxuara Kellner & Campos 1988W13, U03
                      |--T. cristataFMB03
                      |--T. deliradamusW13
                      |--T. leonardii Kellner & Campos 1994K03
                      `--T. longicristatus Kellner & Campos 1989B93

*Lonchodectes compressirostris (Owen 1851) (n. d.) [=Pterodactylus compressirostris, Ornithocheirus compressirostris, Ornithochirus (l. c.) compressirostris, Ornthocheirus (l. c.) compressirostris]RK13

Palaeornis cliftii Mantell 1844 (n. d.) [=Ornithocheirus clifti, Ornithochirus (l. c.) clifti; incl. Pterodactylus ornis Owen 1861, Pt. sylvestris Owen 1859]RK13

*Type species of generic name indicated

References

[A08] Averianov, A. O. 2008. Superorder Pterosauria. In: Ivakhnenko, M. F., & E. N. Kurochkin (eds) Fossil Vertebrates of Russia and Adjacent Territories. Fossil Reptiles and Birds. Part 1 pp. 319–342. GEOS: Moscow.

[B93] Benton, M. J. 1993. Reptilia. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 681–715. Chapman & Hall: London.

[FMB03] Frey, E., D. M. Martill & M.-C. Buchy. 2003. A new species of tapejarid pterosaur with soft-tissue head crest. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 65–72. The Geological Society: London.

[K03] Kellner, A. W. A. 2003. Pterosaur phylogeny and comments on the evolutionary history of the group. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 105–137. The Geological Society: London.

[KC02] Kellner, A. W. A., & D. de A. Campos. 2002. The function of the cranial crest and jaws of a unique pterosaur from the Early Cretaceous of Brazil. Science 297: 389–392.

[RK13] Rodrigues, T., & A. W. A. Kellner. 2013. Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England. ZooKeys 308: 1–112.

[U03] Unwin, D. M. 2003. On the phylogeny and evolutionary history of pterosaurs. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 139–190. The Geological Society: London.

[W13] Witton, M. P. 2013. Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press: Princeton (New Jersey).

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