Basalhaplogynae

Desert bush spider Diguetia canities, from the National Park Service.
Belongs within: Araneida.
Contains: Scytodoidea, Pholcidae, Segestriidae, Dysderidae, Orsolobidae, Oonopidae.

The Basalhaplogynae are a clade of spiders in which the chelicerae are fused at the base and provided with a lamina; members of the clade also have the tegulum and subtegulum fused and the spinnerets lack tartipores (Jocqué & Dippenaar-Schoeman 2007).

<==Basalhaplogynae [Haplogynae, Pholcoidea] W19
|–Filistatidae [Filistatomorpha] JD-S07
|    |  i. s.: Misionella P05
|    |           |–M. didicostae Penney 2005 P05
|    |           `–M. mendensis R14
|    |         Microfilistata Zohnstein 1990 G95
|    |–Filistatinae G95
|    |    |  i. s.: Zaituna Lehtinen 1967 G95
|    |    |–Sahastata Benoit 1968 G95
|    |    |    |–S. nigra G95
|    |    |    `–S. sabaea Brignoli 1982
|    |    `–+–Kukulcania Lehtinen 1967 G95
|    |       |    `–K. hibernalis G95
|    |       `–Filistata Latreille 1810 G95, JD-S07
|    |            |–F. hibernalis B06
|    |            `–F. insidiatrix (Forskål 1775) MG03
|    `–Prithinae G95
|         |  i. s.: Malalistata Mello-Leitão 1946 G95
|         |         Tricalamus Wang 1987 G95
|         |–+–Filistatinella Gertsch & Ivie 1936 G95
|         |  `–+–Filistatoides P.-Cambridge 1899 G95
|         |     `–Pikelinia Mello-Leitão 1946 G95
|         |          `–P. tambilloi R14
|         `–+–+–Andoharano Lehtinen 1967 G95
|            |  `–Afrofilistata Benoit 1968 G95
|            `–+–Pritha Lehtinen 1967 G95, MG03
|               |    |–*P. nana (Simon 1868) MG03
|               |    `–P. crosbyi (Spassky 1938) [=Filistata crosbyi] MG03
|               `–+–Yardiella Gray 1994 G95
|                  `–Wandella Gray 1994 G95
|                       |–W. orana G95
|                       `–W. waldockae MC13
`–Synspermiata [Scytodiformes] W19
|–+–Scytodoidea JD-S07
|  `–+–Pholcidae JD-S07
|     `–+–Plectreuridae JD-S07
|        |    |–Kibramoa Chamberlin 1924 JD-S07
|        |    `–Plectreurys Simon 1893 JD-S07
|        |         `–P. tristis Simon 1893 HF08
|        `–Diguetidae JD-S07
|             |–Segestrioides Keyserling 1883 JD-S07
|             |    `–S. bicolor JD-S07
|             `–Diguetia Simon 1895 JD-S07
|                  |–D. canities JD-S07
|                  `–D. signata Gertsch 1958 HF08
`–+–+–Dysderoidea JD-S07
|  |    |  i. s.: Trogloraptor [Trogloraptoridae] R14
|  |    |–Segestriidae JD-S07
|  |    `–+–Dysderidae JD-S07
|  |       `–+–Orsolobidae JD-S07
|  |          `–Oonopidae JD-S07
|  `–Tetrablemmidae JD-S07
|       |–Shearella JD-S07
|       |–Afroblemma JD-S07
|       |–Tetrablemma P.-Cambridge 1873 [Tetrablemminae] JD-S07
|       |–Paculla [Pacullinae] JD-S07
|       `–Monoblemma spinosum Wunderlich 1988 S93
`–Caponiidae JD-S07
|–Caponia Simon 1887 [Caponiinae] JD-S07
|–Calponia Platnick 1993 JD-S07
|–Diploglena Purcell 1904 JD-S07
|–Nopsides Chamberlin 1924 JD-S07
|–Notnops Platnick 1994 JD-S07
|–Orthonops Chamberlin 1924 JD-S07
|–Taintnops Platnick 1994 JD-S07
|–Tarsonops Platnick 1994 JD-S07
|–Tisentnops Platnick 1994 JD-S07
|–Caponina Simon 1887 JD-S07
|    `–C. notabilis (Mello-Leitão 1939) FM11
`–Nops MacLeay 1839 [Nopinae] JD-S07
|–N. lobatus Wunderlich 1988 S93
`–N. segmentatus Wunderlich 1988 S93

*Type species of generic name indicated

REFERENCES

[B06] Banks, N. 1906. Arachnida from the Bahamas. Bulletin of the American Museum of Natural History 22: 185–189.

[FM11] Freire-Jr, G. de B., & P. C. Motta. 2011. Effects of experimental fire regimes on the abundance and diversity of cursorial arachnids of Brazilian savannah (cerrado biome). Journal of Arachnology 39 (2): 263–272.

[G95] Gray, M. R. 1995. Morphology and relationships within the spider family Filistatidae (Araneae: Araneomorphae). Records of the Western Australian Museum Supplement 52: 79–89.

[HF08] Huber, B. A., & N. Fleckenstein. 2008. Comb-hairs on the fourth tarsi in pholcid spiders. Journal of Arachnology 36 (2): 232–240.

[JD-S07] Jocqué, R., & A. S. Dippenaar-Schoeman. 2007. Spider Families of the World. Royal Museum for Central Africa: Tervuren (Belgium).

[MC13] Majer, J. D., S. K. Callan, K. Edwards, N. R. Gunawardene & C. K. Taylor. 2013. Baseline survey of the terrestrial invertebrate fauna of Barrow Island. Records of the Western Australian Museum Supplement 83: 13–112.

[MG03] Marusik, Yu. M., & E. F. Guseinov. 2003. Spiders (Arachnida: Aranei) of Azerbaijan. 1. New family and genus records. Arthropoda Selecta 12 (1): 29–46.

[P05] Penney, D. 2005. Fossil blood droplets in Miocene Dominican amber yield clues to speed and direction of resin secretion. Palaeontology 48 (5): 925–927.

[R14] Ramírez, M. J. 2014. The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History 390: 1–374.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[W19] Wunderlich, J. 2019. What is a spider? Cretaceous fossils modify strongly phylogenetics as well as diagnoses of families, superfamilies and even suborders of spiders (Araneida) and other arthropods. Beiträge zur Araneologie 12: 1–32.

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