Basalhaplogynae

Desert bush spider Diguetia canities, from the National Park Service.

Belongs within: Araneida.
Contains: Prithinae, Scytodoidea, Tetrablemmidae, Pholcidae, Segestriidae, Oonopidae, Dysderidae, Orsolobidae.

The Basalhaplogynae are a clade of spiders in which the chelicerae are fused at the base and provided with a lamina; members of the clade also have the tegulum and subtegulum fused and the spinnerets lack tartipores (Jocqué & Dippenaar-Schoeman 2007).

<==Basalhaplogynae [Haplogynae, Pholcoidea]W19
    |--Filistatidae [Filistatomorpha]JD-S07
    |    |  i. s.: Microfilistata Zohnstein 1990G95
    |    |--FilistatinaeG95
    |    |    |--Sahastata Benoit 1968G95
    |    |    |    |--S. nigraG95
    |    |    |    `--S. sabaea Brignoli 1982G95
    |    |    `--+--Kukulcania Lehtinen 1967G95
    |    |       |    `--K. hibernalisG95
    |    |       `--+--Zaituna Lehtinen 1967WC17, G95
    |    |          |    `--Z. beshkenticaWC17
    |    |          `--Filistata Latreille 1810G95, JD-S07
    |    |               |--F. gibsonhilli Savory 1943JG19
    |    |               |--F. hibernalisB06
    |    |               |--F. insidiatrix (Forskål 1775)MG03
    |    |               `--F. truncata Risso 1826R26
    |    `--PrithinaeG95
    `--Synspermiata [Scytodiformes]W19
         |--+--ScytodoideaJD-S07
         |  `--+--+--TetrablemmidaeWC17
         |     |  `--+--PlectreuridaeWC17
         |     |     |    |--Kibramoa Chamberlin 1924JD-S07
         |     |     |    |    `--K. madronaWC17
         |     |     |    `--Plectreurys Simon 1893JD-S07
         |     |     |         |--P. desertaWC17
         |     |     |         `--P. tristis Simon 1893HF08
         |     |     `--+--DiguetidaeWC17
         |     |        |    |--Segestrioides Keyserling 1883JD-S07
         |     |        |    |    |--S. bicolorJD-S07
         |     |        |    |    `--S. tofoWC17
         |     |        |    `--Diguetia Simon 1895JD-S07
         |     |        |         |  i. s.: D. signata Gertsch 1958HF08
         |     |        |         |--D. canitiesWC17
         |     |        |         `--+--D. catamarquensisWC17
         |     |        |            `--D. mojaveaWC17
         |     |        `--Pacullidae [Pacullinae]WC17
         |     |             |  i. s.: SabahyaWC17
         |     |             |--Perania nasutaWC17
         |     |             `--+--LamaniaWC17
         |     |                `--Paculla Simon 1887WC17
         |     |                     `--*P. granulosa (Thorell 1881) [=Phaedima granulosa]WC17
         |     `--PholcidaeWC17
         `--+--DysderoideaJD-S07
            |    |--SegestriidaeJD-S07
            |    `--+--OonopidaeWC17
            |       `--+--DysderidaeWC17
            |          `--OrsolobidaeWC17
            `--+--Trogloraptor [Trogloraptoridae]WC17
               |    `--T. marchingtoniWC17
               `--CaponiidaeWC17
                    |--Caponia Simon 1887 [Caponiinae]JD-S07
                    |--Calponia Platnick 1993JD-S07
                    |--Diploglena Purcell 1904JD-S07
                    |--Nopsides Chamberlin 1924JD-S07
                    |--Notnops Platnick 1994JD-S07
                    |--Orthonops Chamberlin 1924JD-S07
                    |--Taintnops Platnick 1994JD-S07
                    |--Tarsonops Platnick 1994JD-S07
                    |--Tisentnops Platnick 1994JD-S07
                    |--Caponina Simon 1887JD-S07
                    |    `--C. notabilis (Mello-Leitão 1939)FM11
                    `--Nops MacLeay 1839 [Nopinae]JD-S07
                         |--N. lobatus Wunderlich 1988S93
                         `--N. segmentatus Wunderlich 1988S93

*Type species of generic name indicated

References

[B06] Banks, N. 1906. Arachnida from the Bahamas. Bulletin of the American Museum of Natural History 22: 185–189.

[FM11] Freire-Jr, G. de B., & P. C. Motta. 2011. Effects of experimental fire regimes on the abundance and diversity of cursorial arachnids of Brazilian savannah (cerrado biome). Journal of Arachnology 39 (2): 263–272.

[G95] Gray, M. R. 1995. Morphology and relationships within the spider family Filistatidae (Araneae: Araneomorphae). Records of the Western Australian Museum Supplement 52: 79–89.

[HF08] Huber, B. A., & N. Fleckenstein. 2008. Comb-hairs on the fourth tarsi in pholcid spiders. Journal of Arachnology 36 (2): 232–240.

[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.

[JD-S07] Jocqué, R., & A. S. Dippenaar-Schoeman. 2007. Spider Families of the World. Royal Museum for Central Africa: Tervuren (Belgium).

[MG03] Marusik, Yu. M., & E. F. Guseinov. 2003. Spiders (Arachnida: Aranei) of Azerbaijan. 1. New family and genus records. Arthropoda Selecta 12 (1): 29–46.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[WC17] Wheeler, W. C., J. A. Coddington, L. M. Crowley, D. Dimitrov, P. A. Goloboff, C. E. Griswold, G. Hormiga, L. Prendini, M. J. Ramírez, P. Sierwald, L. Almeida-Silva, F. Alvarez-Padilla, M. A. Arnedo, L. R. Benavides Silva, S. P. Benjamin, J. E. Bond, C. J. Grismado, E. Hasan, M. Hedin, M. A. Izquierdo, F. M. Labarque, J. Ledford, L. Lopardo, W. P. Maddison, J. A. Miller, L. N. Piacentini, N. I. Platnick, D. Polotow, D. Silva-Dávila, N. Scharff, T. Szűts, D. Ubick, C. J. Vink, H. M. Wood & J. Zhang. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33: 574–616.

[W19] Wunderlich, J. 2019. What is a spider? Cretaceous fossils modify strongly phylogenetics as well as diagnoses of families, superfamilies and even suborders of spiders (Araneida) and other arthropods. Beiträge zur Araneologie 12: 1–32.

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