Batagurinae

 Female Chinese striped-neck turtle Ocadia sinensis, photographed by Tim McCormack.

Belongs within: Pantestuguria.

The Batagurinae are a clade of freshwater turtles found in Asia.

Synapomorphies (from Gaffney & Meylan 1988): Fissura ethmoidalis oval and broad; frontals sometimes excluded from orbital rim by prefrontal-postorbital contact; foramen palatinum posterius small, excluded from pterygoid.

<==Batagurinae
    |--+--Orlitia borneensis (Gray 1873)GM88, JPG04
    |  `--Siebenrockiella crassicollis (Gray 1831)GM88, JPG04
    `--+--+--Geoclemys hamiltonii (Gray 1831)GM88, JPG04 [=Damonia hamiltoniUSDI77]
       |  `--+--Malayemys subtrijuga (Schlegel & Müller 1844)GM88, JPG04
       |     `--ChinemysGM88
       |          |--C. megalocephalaCP03
       |          |--C. nigricansCP03
       |          `--C. reevesiCP03
       `--+--Hieremys annandalii (Boulenger 1903)GM88, JPG04
          `--+--Ocadia sinensisGM88, CP03
             `--+--+--Batagur baska (Gray 1831)GM88, JPG04 [=Emys baskaJPG04]
                |  `--Callagur borneoensis (Schlegel & Müller 1844)GM88, JPG04
                |--+--Hardella thurjii (Gray 1831)GM88, JPG04
                |  `--MoreniaGM88
                |       |--M. ocellata (Duméril & Bibron 1835)JPG04
                |       `--M. petersi (Anderson 1879)JPG04
                `--KachugaGM88
                     |--K. dhongokha (Gray 1834)JPG04
                     |--K. kachuga (Gray 1831)JPG04
                     |--K. smithiiCP03
                     |--K. tectaCP03
                     |--K. tentoriaCP03
                     `--K. trivittata (Duméril & Bibron 1835)JPG04
Hybrids: Ocadia glyphistoma McCord & Iverson 1994 [O. sinensis × Mauremys annamensis]D03
         Ocadia philippeni McCord & Iverson 1992 [O. sinensis × Cuora trifasciata]D03

*Type species of generic name indicated

References

[CP03] Claude, J., E. Paradis, H. Tong & J.-C. Auffray. 2003. A geometric morphometric assessment of the effects of environment and cladogenesis on the evolution of the turtle shell. Biological Journal of the Linnean Society 79: 485–501.

[D03] Dalton, R. 2003. Mock turtles. Nature 423: 219–220.

[GM88] Gaffney, E. S., & P. A. Meylan. 1988. A phylogeny of turtles. In: Benton, M. J. (ed.) The Phylogeny and Classification of the Tetrapods vol. 1. Amphibians, Reptiles, Birds pp. 157–219. Clarendon Press: Oxford.

[JPG04] Joyce, W. G., J. F. Parham & J. A. Gauthier. 2004. Developing a protocol for the conversion of rank-based taxon names to phylogenetically defined clade names, as exemplified by turtles. Journal of Paleontology 78 (5): 989–1013.

[USDI77] USDI (United States Department of the Interior). 1977. Endangered and threatened wildlife and plants—republication of list of species. Federal Register 42: 36420–36431.

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