Bothriuridae

Black scorpion Bothriurus bonariensis, photographed by Gustavo Schmidt.

Belongs within: Iurida.

The Bothriuridae is a group of scorpions found in Australia and South America characterised by legs with two pedal spurs, a vertically compressed sternum, and pedipalps with chelal trichobothrium Et2 on the ventral surface and ib at the extreme base of the fixed finger or on the palm.

Bothriuridae [Bothriuroidae, Bothriuroidea]
|--VachonianinaeK77
|--Brachistosternus Pocock 1893OA02b [BrachistosterninaeK77]
| | i. s.: B. andinus Chamberlin 1916OA02b
| | B. ehrenbergii (Gervais 1841)OA02b [incl. B. ehrenbergi var. politusAO02]
| | B. ferrugineus (Thorell 1877)AO02
| | B. holmbergi Carbonell 1923OA02b
| | B. peruvianus Piza 1974OA02b
| `--B. (Leptosternus Maury 1973)OA02b
| |--B. (L.) castroi Mello-Leitão 1940OA02b
| |--B. (L.) intermedius Lönnberg 1902OA02b
| |--B. (L.) montanus Roig Alsina 1977OA02b
| |--B. (L.) quiscapata Ochoa & Acosta 2002OA02b
| |--B. (L.) titicaca Ochoa & Acosta 2002OA02b
| `--B. (L.) weijenberghii (Thorell 1876)OA02b
|--LisposomaVP08 [LipsominaeK77]
| `--L. josehermana Lamoral 1979VP08
`--Bothriurinae [Cercophoninae]K77
|--Cercophonius Peters 1861 [incl. Acanthochirus Peters 1861]K77
| `--*C. squama (Gervais 1844) (see below for synonymy)K77
|--Urophonius Pocock 1893K77
| |--U. brachycentrus (Thorell 1877)K77
| `--U. jeheringi Pocock 1893K77
|--Timogenes Simon 1880K77
| |--T. dorbignyi (Guérin-Méneville 1843) [=Scorpio dorbignyi]AO02
| |--T. elegans (Mello-Leitão 1931)AO02
| `--T. haplochirus Maury & Roig Alsina 1977CFS14
`--Bothriurus Peters 1861CFS14
|--B. araguayae Vellard 1934FM11
|--B. asper Pocock 1893 [=B. bonariensis asper]AO02
|--B. bocki Kraepelin 1911AO02
|--B. bonariensis (Koch 1843)AO02
|--B. bucherli (San Martin 1965)K77
|--B. burmeisteri Kraepelin 1894CFS14
|--B. chacoensis Maury & Acosta 1993AO02
|--B. chilensis (n. d.) [=Scorpio chilensis]P00
|--B. dumayiAO02
|--B. flavidus (Maury 1971)K77
|--B. inermis Maury 1981AO02
|--B. maculatus Kraepelin 1911 [=B. bonariensis var. maculata]AO02
|--B. olaen Acosta 1997AO02
|--B. prospicuusA02
|--B. rochensisK77
|--B. signatusP00
|--B. trivittatus Werner 1939AO02
`--B. vittatus Guérin-Méneville 1838AO02

Bothriuridae incertae sedis:
Orobothriurus Maury 1976OA02a
|--O. alticola (Pocock 1899)OA02a
|--O. atiquipa Ochoa & Acosta 2002OA02a
|--O. calchaqui Ochoa et al. 2011CFS14
|--O. compagnucci Ochoa et al. 2011CFS14
|--O. curvidigitus (Kraepelin 1911)OA02a [incl. Bothriurus lampei Werner 1916J98]
|--O. famatina Acosta & Ochoa 2001CFS14
|--O. grismadoi Ojanguren-Affilastro et al. 2009CSF14
|--O. iskay Acosta & Ochoa 2001AO02
|--O. paessleri (Kraepelin 1911)OA02a
`--O. wawita Acosta & Ochoa 2000AO02
Tehuankea Cekalovic 1973K77
BrandbergiaVP08

*Cercophonius squama (Gervais 1844) [=Scorpio (Telegonus) squama; incl. C. granulosus Kraepelin 1908, C. kershawi Glauert 1930, C. michaelseni Kraepelin 1908, C. sulcatus Kraepelin 1908, *Acanthochirus testudinarius Peters 1861]K77

*Type species of generic name indicated

References

[A02] Acosta, L. E. 2002. Patrones zoogeográficos de los Opiliones Argentinos (Arachnida: Opiliones). Revista Ibérica de Aracnología 6: 69–84.

[AO02] Acosta, L. E., & J. A. Ochoa. 2002. Lista de los escorpiones bolivianos (Chelicerata: Scorpiones), con notas sobre su distribución. Revista de la Sociedad Entomológica Argentina 61 (3–4): 15–23.

[CSF14] Campón, F. F., S. L. Silnik & L. A. Fedeli. 2014. Scorpion diversity of the Central Andes in Argentina. Journal of Arachnology 42 (2): 163–169.

[FM11] Freire-Jr, G. de B., & P. C. Motta. 2011. Effects of experimental fire regimes on the abundance and diversity of cursorial arachnids of Brazilian savannah (cerrado biome). Journal of Arachnology 39 (2): 263–272.

[J98] Jäger, P. 1998. Das Typenmaterial der Spinnentiere (Arachnida: Acari, Amblypygi, Araneae, Opiliones, Pseudoscorpiones, Scorpiones, Uropygi) aus dem Museum Wiesbaden. Jahrbuecher des Nassauischen Vereins fuer Naturkunde 119: 81–91.

[K77] Koch, L. E. 1977. The taxonomy, geographic distribution and evolutionary radiation of Australo-Papuan scorpions. Records of the Western Australian Museum 5 (2): 83–367.

[OA02a] Ochoa, J. A., & L. E. Acosta. 2002a. Orobothriurus atiquipa, a new bothriurid species (Scorpiones) from Lomas in southern Peru. Journal of Arachnology 30: 98–103.

[OA02b] Ochoa, J. A., & L. E. Acosta. 2002b. Two new Andean species of Brachistosternus Pocock (Scorpiones: Bothriuridae). Euscorpius 2: 1–13.

[P00] Pocock, R. I. 1900. Some new or little-known Neotropical scorpions in the British Museum. Annals and Magazine of Natural History, series 7, 5: 469–478.

[VP08] Volschenk, E. S., & L. Prendini. 2008. Aops oncodactylus, gen. et sp. nov., the first troglobitic urodacid (Urodacidae: Scorpiones), with a re-assessment of cavernicolous, troglobitic and troglomorphic scorpions. Invertebrate Systematics 22: 235–257.

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