Liochthonius proprinquus, from here.

Belongs within: Oribatida.
Contains: Liochthonius (Liochthonius), Brachychthonius, Sellnickochthonius.

Liochthonius is an cosmopolitan genus of unornamented brachychthoniid mites in which the adanal plates are fused to form a U-shaped plate surrounding the anal plate (Balogh & Balogh 1992).

Brachychthoniidae [Brachychthonioidea]SC19
    |--Liochthonius Hammen 1959SC19, S04
    |   |--L. (Liochthonius)S04
    |   `--L. (Afroliochthonius Mahunka 1995)S04
    |        `--L. (*A.) reductus Mahunka 1995S04
    `--+--Neoliochthonius Lee 1982SC19, S04 (see below for synonymy)
       |    |--*N. piliforus (Frosslund 1942) [=Brachychthonius piliforus]S04
       |    |--N. globuliferus (Strenzke 1951)S04 [=Brachychthonius globuliferusS04, Paraliochthonius globuliferusBB92]
       |    `--N. occultus (Niedbała 1971) [=Liochthonius occultus]S04
       `--+--Eobrachychthonius Jacot 1936SC19, S04
          |    |--E. latior (Berlese 1910) [=Brachychthonius latior]S04
          |    |    |--E. l. latiorS04 (see below for synonymy)
          |    |    `--E. l. strictior Lombardini 1963 (n. d.)S04
          |    |--E. borealis Forsslund 1942S04
          |    |--E. longisetosus Csiszár 1960S04
          |    |--E. mooseri (Hammen 1951) [=Brachychthonius mooseri]S04
          |    |--E. oudemansi Hammer 1952 [incl. E. argentinensis Hammer 1958]S04
          |    |--E. setus Sergienko 1992S04
          |    `--E. similis Mahunka 1979S04
          `--+--‘Sellnickochthonius’ immaculatus (Forsslund 1942)SC19, S04 (see below for synonymy)
Brachychthoniidae incertae sedis:
  Papillochthonius Gil-Martín, Subías & Arillo 1992S04
    `--*P. astatus Gil-Martín, Subías & Arillo 1992S04
  Neobrachychthonius Moritz 1976S04
    |--*N. marginatus (Forsslund 1942) [=Brachychthonius marginatus; incl. B. neosimplex Schweizer 1956]S04
    `--N. magnus Moritz 1976 [=N. marginatus magnus]S04
  Synchthonius Hammen 1952S04
    |--S. crenulatus (Jacot 1938) [=Brachychochthonius crenulatus; incl. *S. boschmai Hammen 1952]S04
    `--S. elegans Forsslund 1957S04
  Poecilochthonius Balogh 1943S04 [=PoecilochtoniusCH98]
    |--*P. italicus (Berlese 1910) [=Brachychthonius brevis italicus]S04
    |--P. parallelus (Womersley 1945) [=Brachychthonius parallelus]S04
    `--P. spiciger (Berlese 1910) [=Brachychthonius brevis spiciger; incl. B. rapoporti Balogh & Mahunka 1968]S04
  Mixochthonius Niedbała 1972S04
    |--*M. pilosetosus (Forsslund 1942) [=Brachychthonius pilosetosus]S04
    |--M. concavus (Chinone 1974) [=Liochthonius concavus]S04
    |--M. laticepsNP91
    `--M. moritzi Evison 1981S04
  Verachthonius Moritz 1976S04
    |--*V. laticeps (Strenzke 1951) [=Brachychthonius laticeps]S04
    |--V. congruus Moritz 1976S04
    |--V. diversus Moritz 1976S04
    |--V. montanus (Hammer 1952) [=Eobrachychthonius montanus]S04
    |--V. moritzi Lee 1982S04
    `--V. pseudolaticeps (Subías 1977) [=Liochthonius pseudolaticeps]S04

Eobrachychthonius latior latior (Berlese 1910)S04 [incl. Brachychthonius brevis glabra Thor 1930S04, E. grandis Sellnick 1944S04, B. grandisH44, *E. sexnotatus Jacot 1936S04]

Neoliochthonius Lee 1982SC19, S04 [incl. Moritzichthonius Pereda & Iturrondobeitia 1990S04, Paraliochthonius Moritz 1976 non Beier 1956S04]

‘Sellnickochthonius’ immaculatus (Forsslund 1942)SC19, S04 [=Brachychochthonius immaculatusS04; incl. Brachychochthonius arcticus Hammer 1952S04, Brachychochthonius berlesei erosus Jacot 1938S04, Brachychthonius gygeri Bader 1963S04, Brachychthonius obscurus Krivolutsky 1966S04, Brachychthonius seminornatus Evans 1952S04, Brachychthonius striatus Willmann 1956S04]

*Type species of generic name indicated


[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[H44] Hammer, M. 1944. Studies on the oribatids and collemboles of Greenland. Meddelelser om Grønland 141 (3): 1–210.

[NP91] Norton, R. A., & S. C. Palmer. 1991. The distribution, mechanisms and evolutionary significance of parthenogenesis in oribatid mites. In: Schuster, R., & P. W. Murphy (eds) The Acari: Reproduction, development and life-history strategies pp. 107–136. Chapman & Hall: London.

[SC19] Schaefer, I., & T. Caruso. 2019. Oribatid mites show that soil food web complexity and close aboveground-belowground linkages emerged in the early Paleozoic. Communications Biology 2: 387.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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