Brachystigmata

Male (above) and female
Chlorogomphus magnificus, copyright Wildan Ardani.

Belongs within: Cavilabiata.
Contains: Paneurypalpidomorpha.

The Brachystigmata are a clade of dragonflies that mostly have fairly short pterostigmata in the wings (Bechly & Ueda 2002).

Synapomorphies (from Bechly & Ueda 2002): Wings with relatively short pterostigmata that cover only 1–3 complete cells (reversed in Libellulinae); “gaff” (= basal CuA before its branching) in hind wing strongly prolonged and very straight in the groundplan (sigmoidal in Eurypalpidomorpha); area between MP and CuA basally widened with more than one row of cells (reversed in some subclades); terminal branch of CuAa secondarily branched on CuA (inapplicable whe CuAa is secondarily unbranched); RP3/4 and MA strictly parallel up to the hind margin; area between RP2 and IR2 distally distinctly widened, with more than one cell row in the distal half (sometimes reversed).

<==Brachystigmata BU02
|–Nannogomphidae BU02
`–Paucipostnodalia BU02
|–Prohemeroscopus Bechly et al. 1998 [Prohemeroscopidae] BU02
|    |–*P. jurassicus BU02
|    `–P. kuehnapfeli BU02
`–Eubrachystigmata BU02
|–Hemeroscopus R02 [Hemeroscopidae BU02]
|    `–H. baissicus Pritykina 1977 R02
`–Neobrachystigmata BU02
|–Paneurypalpidomorpha BU02
`–Chlorogomphida BU02
|–Araripechlorogomphus Bechly & Ueda 2002 [Araripechlorogomphidae] BU02
|    `–*A. muratai Bechly & Ueda 2002 BU02
`–Chlorogomphoidea BU02
|–Chloropetaliidae [Chloropetaliinae] BU02
`–Chlorogomphus SL89 [Chlorogomphinae BU02]
|–C. brevistigma I92
|    |–C. b. brevistigma I92
|    `–C. b. okinawensis Ishida 1964 I92
|–C. brunneus Oguma 1926 I92
|    |–C. b. brunneus I92
|    |–C. b. costalis I92
|    `–C. b. keramensis Asahina 1972 I92
|–C. hyalinus SL89
|–C. iriomotensis Ishida 1972 I92
|–C. magnificus SL89
`–C. papilio RD77

*Type species of generic name indicated

REFERENCES

[BU02] Bechly, G., & K. Ueda. 2002. The first fossil record and first New World record for the dragonfly clade Chlorogomphida (Insecta: Odonata: Anisoptera: Araripechlorogomphidae n. fam.) from the Crato Limestone (Lower Cretaceous, Brazil). Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 328: 1–11.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[R02] Rasnitsyn, A. P. 2002. Special features of the study of fossil insects. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 8–12. Kluwer Academic Publishers: Dordrecht.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[SL89] Selys Longchamps, E. de. 1889. Odonates de Sumatra comprenant les espèces recueillies à Pulo Nias par M. le D.r E. Modigliani. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 444–484.

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