Xanthocaecilius quillayute, photographed by Dann Thombs.

Belongs within: Caeciliusetae.
Contains: Enderleinella, Paracaecilius, Fuelleborniella, Coryphacini, Austrocaeciliini, Maoripsocini, Caecilius, Dypsocus.

The Caeciliusidae are a group of barklice with forewing veins bearing setae on the dorsal surface only. They are found living on vegetation, most commonly on living trees (New & Lienhard 2007). The Australian species Aphyopsocus prolixus is distinguished by having the basal flagellar segment of the antenna enlarged (Schmidt & New 2008).

Characters (from Mockford 1993): Inner clypeal shelf variable. Distal inner labral sensilla five to seven. Lacinial tip variable: broad and flat, broad and denticulate, slender and bicuspid, or slender and bilobed. Labral stylets present or absent. Maxillary palpus with mx4 shorter than mx2. Setae of forewings slanting distad, relatively short, usually in one rank except on costa. Setae on veins in distal half of forewing only on dorsal surface. Hindwing margin mostly ciliated except basal two-thirds of front margin. Spine of free margin of paraproct small or absent. Endophallus with median bulb, usually not divided on midline, ovipositor valvulae with v3 a sclerotic bulge from base of v2.

<==Caeciliusidae [Caeciliini]
    |--Aphyopsocus Smithers 1982 [Aphyopsocinae]SN08
    |    `--*A. prolixus Smithers 1982SN08
    |    |--EnderleinellaNL07
    |    `--ParacaeciliusSN08
    |--Amphicaecilius Li 2002 [Amphicaeciliinae]L02
    |    |--*A. pulcherrimus Li 2002L02
    |    |--A. daqingshanicus Li 2002L02
    |    |--A. floribundus Li 2002L02
    |    `--A. pulchellus Li 2002L02
    |--Fuelleborniellinae [Fuellebornelliini]NL07
    |    |--FuelleborniellaS95
    |    `--YpsiloneuraB55
    |         |--Y. kirkpatrickiB55
    |         `--Y. monostyla Badonnel 1955B55
    |--Isophanes Banks 1937 [Isophaneinae]L02
    |    |--*I. decipiens Banks 1937L02
    |    |--I. angolensis Badonnel 1955B55
    |    |--I. cyclostigma Li 1993L02
    |    |--I. fengduensis Li 1997L02
    |    |--I. lutea Li 2002L02
    |    |--I. orientalis Garcia Aldrete 1999 [incl. I. sinensis Li 1993 non New 1991]L02
    |    |--I. oxyostigma Li 1993L02
    |    |--I. platytigma Li 1997L02
    |    |--I. sinensis New 1991L02
    |    `--I. yangi Li 1993L02
    `--Caeciliusinae [Dypsocinae]SN08
         |  i. s.: Disialacaecilia Li 2002L02 [=DisiacaeciliaL03, DisialacaeciliusL03]
         |           `--*D. ningxiaensis Li 2002L02
         |         Parvialacaecilia Li 2002L02
         |           `--*P. hebeiensis Li 2002L02 [=P. hedeiensisL03]
         |         Anoculaticaeca Li 1997L03, L02 [=Anoculaticaecilius (l. c.)L03]
         |           `--*A. chuanshaanica Li 1997L02, L03, L02 [=Anoculaticaecilius chuanshaanicus (l. c.)L03]
         |         Licaecilius Lienhard 2003L03 [=Neocaecilius Li 2002 non Li 2000L02]
         |           |--*L. triradiatus (Li 2002) [=*Neocaecilius triradiatus]L03
         |           `--L. mangshiensis (Li 2002) [=Neocaecilius mangshiensis]L03
         |         Bivalvicaecilia Li 2002L02 [=BivalvicaeciliusL03]
         |           |--*B. longiansa Li 2002L02
         |           `--B. abbreviata (Li 1993) [=Caecilius abbreviatus, Asiocaecilius abbreviatus]L02
         |         Phymocaecilius Li 2002L02 [=PhtmocaeciliusL03, PhymocaecilusL03, PhymocaeiliusL03]
         |           |--*P. subulosus Li 2002L02
         |           |--P. fortis (Li 1995) [=Caecilius fortis, Velenzuala forti]L02
         |           |--P. fuscifascus Li 2002L02
         |           |--P. guizhouensis Li 2002L02
         |           `--P. puniceifascus Li 2002L02
         |    |--CaeciliusNL07
         |    `--DypsocusNL07
         |    |--Bassocaecilius Schmidt & New 2008SN08
         |    |    `--*B. rawlinsoni Schmidt & New 2008SN08
         |    `--Clinocaecilius Schmidt & New 2008SN08
         |         `--*C. edwardsi Schmidt & New 2008SN08
              |--Nothocaecilius Schmidt & New 2008SN08
              |    `--*N. thomasi Schmidt & New 2008SN08
              `--Tasmanocaecilius Schmidt & New 2008SN08
                   `--*T. truchanasi Schmidt & New 2008SN08
Caeciliusidae incertae sedis:
  Xanthocaecilius Mockford 1989M93
    |--X. quillayute (Chapman 1930) [=Caecilius quillayute]M93
    `--X. sommermanae (Mockford 1955) [=Caecilius sommermanae]M93
  Orocaecilius Mockford 1999L02
  Lacroixiella Badonnel 1943L98
    `--*L. martini (Lacroix 1919) [=Caecilius martini]L98
  Eopsocites Hong 2002H02
    `--*E. fushunensis Hong 2002H02
  Stenopterites Hong 2002H02
    |--*S. guchengziensis Hong 2002H02
    `--S. longus Hong 2002H02
  Fushunopsocus Hong 2002H02
    `--*F. latus Hong 2002H02
  Palaeopsocus Kolbe 1883H02
  Ptenolasia Enderlei 1991H02
  Liaoxipsocus Hong 1983H02
    `--L. beipiaoensis (Hong 1983)L02
  Beipiaopsocus Hong 1983H02
    `--B. divaricatus (Hong 1983)L02
  Huaxiapsocus Hong 1983H02
    `--H. parvus (Hong 1983)L02
  Psocites Hong 1983 [incl. Parapsocus Hong 1983]H02
    `--P. pectinatus (Hong 1992)L02

*Type species of generic name indicated


[B55] Badonnel, A. 1955. Psocoptères de l’Angola. Diamang Publicações Culturais 26: 11–267.

[H02] Hong Y. 2002. Amber Insect of China. Beijing Scientific and Technological Publishing House.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529–561.

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