Calopterygoidea

 Male yellow bush dart Psilocnemis marginipes, copyright Jee & Rani Nature Photography.

Belongs within: Zygoptera.
Contains: Rhinocypha, Agrion.

The Calopterygoidea are a group of damselflies characterised by broad, often colourful wings. In members of the family Calopterygidae, the wings are densely reticulate with numerous cross- and supplementary longitudinal veins, and have non-petiolate bases. Historically, this family has been recognised as the Agrionidae or Agriidae, a name that has also been used to encompass many, or all, of the damselflies. Modern classifications do not include this family with the type genus Agrion no longer being treated as a valid name.

Other families include the Chlorocyphidae, an Old World tropical group with a narrow head, close eyes, and a produced, snout-like clypeus. The Polythoridae are a South American family in which the wings lack an arculus and have the base of the discoidal cell connected directly to R5 (Richards & Davies 1977).

<==Calopterygoidea [Agrioidea, Calopterygomorpha]
    |--EosagrionidaeFB03
    |--DicteriastidaeWO91
    |--Pseudolestes Kirby 1900K00 [PseudolestidaeRD77]
    |    `--*P. mirabilis Kirby 1900K00
    |--HeliocharitidaeRD77
    |--PolythoridaeWO91
    |    |--Euthore fasciata (Hagen 1853)FB03
    |    `--CoraZP01
    |         |--C. chiribiquete Zloty & Pritchard 2001ZP01
    |         |--C. mundaZP01
    |         `--C. terminalisZP01
    |--EuphaeidaeWO91
    |    |--Epallagites avusRJ93
    |    `--EuphaeaSL89
    |         |--E. aspasiaSL89
    |         |--E. bocki MacLachl. 1880SL89
    |         |--E. variegataSL89
    |         `--E. yayeyamensisI92
    |--Chlorocyphidae [Libellaginidae]FB03
    |    |--Libellago Selys 1840FB03
    |    |--Heliocypha fenestrataTH06
    |    `--RhinocyphaTH06
    |--Calopterygidae [Calopteryginae]W75
    |    |--Psolodesmus mandarinus McLachlan 1870FB03
    |    |--NeurobasisTH06
    |    |    |--N. australis Selys 1897TH06
    |    |    `--N. chinensisTH06
    |    |--MnaisFB03
    |    |    |--M. andersoni McLachlan 1873FB03
    |    |    |--M. nawaiTMS91
    |    |    `--M. pruinosaTMS91
    |    |--Micromerus Rambur 1842SL89, BP02
    |    |    |--M. aurantiacusSL89
    |    |    |--M. snellemaniSL89
    |    |    `--M. sumatranusSL89
    |    |--CalopteryxW75
    |    |    |--C. aequabilisW75
    |    |    |--C. maculataW75
    |    |    |--C. splendensW75
    |    |    `--C. virgoW75
    |    |--VestalisSL89
    |    |    |--V. amaenaSL89
    |    |    |--V. luctuosaSL89
    |    |    |--V. lugens [incl. V. coracina]SL89
    |    |    `--V. melaniaSL89
    |    `--Dysphaea Selys 1853SL89, PN04
    |         |--D. dimidiataSL89
    |         |--D. limbataSL89
    |         |--D. lugensSL89
    |         `--D. sublimbataSL89
    `--Agriidae [Agrionidae, Agrioninae]RD77
         |--Onychargia atrocyanaSL89
         |--Stenobasis oscillansSL89
         |--Allonevra insignisSL89
         |--PodolestesSL89
         |    |--P. chrysopus Selys Longchamps 1889SL89
         |    `--P. orientalisSL89
         |--Trichocnemis membranipes [=Platycnemis membranipes]SL89
         |    |--T. m. membranipesSL89
         |    `--T. m. silentaSL89
         |--DisparonevraSL89
         |    |--D. notostigmaSL89
         |    `--D. verticalisSL89
         |--SympycnaK01
         |    |--S. fuscaK01
         |    `--S. paediscaK01
         |--Erythromma najasK01
         |    |--E. n. najasI92
         |    `--E. n. baicalensis Belyshev 1964I92
         |--AgrionK01
         |--PsilocnemisSL89
         |    |--P. annulataSL89
         |    |    |--P. a. annulataSL89
         |    |    |--P. a. ciliataSL89
         |    |    |--P. a. serapicaSL89
         |    |    `--P. a. subannulataSL89
         |    |--P. imbricataSL89
         |    `--P. marginipes [=Platycnemis marginipes, incl. Pl. lacteola, Psilocnemis striatipes]SL89
         |--Matrona basilaris Selys 1853K00
         |    |--M. b. basilarisI92
         |    |--M. b. japonica Foerster 1897I92
         |    `--M. b. nigripectusI92
         |--Bayadera brevicaudaK00, I92
         |    |--B. b. brevicaudaI92
         |    `--B. b. ishigakiana Asahina 1964I92
         `--Pseudophaea decorata (de Selys 1853) [=Euphaea decorata]K00

*Type species of generic name indicated

References

[BP02] Bologna, M. A., & J. D. Pinto. 2002. The Old World genera of Meloidae (Coleoptera): a key and synopsis. Journal of Natural History 36 (17): 2013–2102.

[FB03] Fleck, G., G. Bechly, X. Martínez-Delclòs, E. Jarzembowski, R. Coram & A. Nel. 2003. Phylogeny and classification of the Stenophlebioptera (Odonata: Epiproctophora). Annales de la Societe Entomologique de France 39 (1): 55–93.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[K00] Kirby, W. F. 1900. On a small collection of Odonata (dragonflies) from Hainan, collected by the late John Whitehead. Annals and Magazine of Natural History, series 7, 5: 530–539, pl. 12.

[K01] Klapálek, F. 1901. Neuropteroidák [Neuropteroiden]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 203–221. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[PN04] Petrulevičius, J. F., & A. Nel. 2004. A new damselfly family from the Upper Palaeocene of Argentina. Palaeontology 47: 109–116.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[SL89] Selys Longchamps, E. de. 1889. Odonates de Sumatra comprenant les espèces recueillies à Pulo Nias par M. le D.r E. Modigliani. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 444–484.

[TH06] Theischinger, G., & J. Hawking. 2006. The Complete Field Guide to Dragonflies of Australia. CSIRO Publishing: Collingwood (Victoria).

[TMS91] Tsurusaki, N., M. Murakami & K. Shimokawa. 1991. Geographic variation of chromosomes in the Japanese harvestman, Gagrellopsis nodulifera, with special reference to a hybrid zone in western Honshu. Zoological Science 8: 265–275.

[W75] Waage, J. K. 1975. Reproductive isolation and the potential for character displacement in the damselflies Calopteryx maculata and C. aequabilis (Odonata: Calopterygidae). Systematic Zoology 24 (1): 24–36.

[WO91] Watson, J. A. L., & A. F. O’Farrell. 1991. Odonata (dragonflies and damselflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 294–310. Melbourne University Press: Carlton (Victoria).

[ZP01] Zloty, J., & G. Pritchard. 2001. Cora chiribiquete spec. nov., a new damselfly species from Colombia (Zygoptera: Polythoridae). Odonatologica 30 (2): 227–232.

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