Cognate land snail Amphidromus cognatus, from here.

Belongs within: Helicoidei.

The Camaeninae are a group of snails found between southern China and the Solomon Islands and northern and eastern Australia. Members of this group are distinguished by an eversible head wart together with a genital system including a long tubular verge and elongated penis complex. Species found in South Australia commonly have the radula modified for continuously scraping particles from rock surface, whereas species from elsewhere have a more typical radula for chopping and cutting plant matter (Solem 1997).

<==Camaeninae [Epiphallogona, Epiphallophora]
    |--Damochlora Iredale 1938S97
    |--Contramelon Iredale 1937S97
    |--Cooperconcha Solem 1992S97
    |--Pseudcupedora Solem 1992S97
    |--Aslintesta Solem 1992S97
    |--Cupedora Iredale 1933S97
    |    `--C. extensa (Iredale 1937)S97
    |--Glyptorhagada Pilsbry 1890S97
    |    `--G. bordaensis (Angas 1880)S97
    |--Offachloritis Iredale 1933S97, SS10
    |    |--*O. dryanderensis (Cox 1872) [=Helix dryanderensis]SS10
    |    `--O. melaniecarlessae Stanisic in Stanisic, Shea et al. 2010SS10
    |--Amphidromus Albers 1850 [Amphidrominae]BR05
    |    |--*A. perversus (Linnaeus 1758) [=Helix perversa]BR17
    |    |--A. cognatus Fulton 1907S97
    |    `--A. formosensis (Adams 1866) [=Bulimus (Amphidromus) formosensis]A66
    `--Camaena Albers 1850BR05
         |--*C. cicatricosa (Müller 1774) [=Helix cicatricosa]BR17
         |--C. bairdi (Adams 1866) [=Helix (Camaena) bairdi]A66
         `--C. succincta (Adams 1866) [=Helix (Camaena) succincta]A66

*Type species of generic name indicated


[A66] Adams, H. 1866. Descriptions of fifteen new species of land and freshwater shells from Formosa, collected by Robert Swinhoe, Esq., consul at Taiwan in that island. Proceedings of the Zoological Society of London 1866: 316–319.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[S97] Solem, A. 1997. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). VII. Taxa from Dampierland through the Nullarbor. Records of the Western Australian Museum Supplement 50: 1461–1906.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

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