Camisia cf. nova, photographed by S. E. Thorpe.

Belongs within: Crotoniidae.

Camisia is a cosmopolitan genus of primarily arboreal oribatid mites, with some species found among mosses on rocks. Specimens of the genus are known from Baltic amber. Camisia is divided between two subgenera distinguished by whether the legs are tridactylous (Camisia) or monodactylous (Ensicamisia).

Characters (from Balogh & Balogh 1992): Bothridial sensillus present. Notogaster with more or less parallel margins, posteriorly widely truncate. Epimeral setal formula 3-1-3-3 or 3-1-2-3. Genital and anal plates of macropyline type, large, occupying entire ventral side posterior to epimeres; nine pairs of genital setae; two pairs of aggenital setae; three pairs of anal setae; three pairs of adanal setae. Legs with one or three tarsal claws.

<==Camisia Heyden 1826 [=Uronothrus Berlese 1913; Camisiinae]S04
    |--C. (Camisia)S04
    |    |  i. s.: C. (C.) abdosensilla Olszanowski & Clayton 2002S04
    |    |         C. (C.) amictus (Sowerby 1806) (n. d.) [=Acarus amictus; incl. Nothrus angulatus Koch 1839]S04
    |    |         C. (C.) flagellata (Berlese 1888) [=Nothrus flagellatus]S04
    |    |         C. (C.) heterospinifer Fujikawa 1998S04
    |    |         C. (C.) orthogonia Olszanowski, Szywilewska & Norton 2001S04
    |    |         C. (C.) tatrica Olszanowski 1994S04
    |    |         C. (C.) umbratilis (Hull 1915) (n. d.) [=Nothrus umbratilis]S04
    |    |         C. (C.) zangherii Lombardini 1963 (n. d.)S04
    |    |         C. hamulifera groupC93
    |    |           |--C. (C.) arcuata Hammer 1961C93
    |    |           |--C. (C.) grymae Colloff 1993C93
    |    |           |--C. (C.) hamulifera Hammer 1961S04
    |    |           `--C. (C.) valeriae Colloff 1993C93
    |    |         C. spinifer groupC93
    |    |           |--C. (C.) biurus (Koch 1839) [=Nothrus biurus]C93
    |    |           |    |--C. b. biurus (see below for synonymy)S04
    |    |           |    `--C. b. amicus Jacot 1938 [=C. segnis amicus]S04
    |    |           `--C. (C.) spinifer (Koch 1835)S04 (see below for synonymy)
    |    |--C. horrida groupC93
    |    |    |--C. (C.) anomia Colloff 1993C93
    |    |    |--C. (C.) australis Hammer 1958C93
    |    |    |--C. (C.) biverrucata (Koch 1839) (see below for synonymy)C93
    |    |    |--C. (C.) dictyna Colloff 1993C93
    |    |    |--C. (C.) horrida (Hermann 1804)S04, SC19, S04 [=Northrus horridus fossilusC93, C. horrida fossilusC93]
    |    |    |--C. (C.) nortoni Colloff 1993C93
    |    |    `--C. (C.) oregonae Colloff 1993C93
    |    `--+--C. segnis groupC93
    |       |    |--*C. (C.) segnis (Hermann 1804)S04, SC19, S04 (see below for synonymy)
    |       |    |--C. (C.) carrolli André 1980C93
    |       |    |--C. (C.) fossilus (Sellnick 1919) (n. d.) (see below for synonymy)C93
    |       |    |--C. (C.) khencensis Hammer 1961C93
    |       |    |--C. (C.) nova Hammer 1966 [=C. segnis var. nova]C93
    |       |    `--C. (C.) tryphosa Colloff 1993C93
    |       `--C. invenusta groupC93
    |            |--C. (C.) foveolata Hammer 1955C93
    |            |--C. (C.) invenusta (Michael 1888)S04, SC19, S04 [=Nothrus invenustusS04]
    |            `--C. (C.) polytricha Piffl 1971C93
    `--C. (Ensicamisia Kunst 1971) [incl. Ivarsia Karppinen & Krivolutsky 1987]S04
         |--C. (*E.) lapponica (Trägardh 1910) [=Nothrus lapponicus; incl. C. labradorica Behan-Pelletier 1978]S04
         |--C. (E.) presbytis Colloff 1993S04
         |--C. (E.) sibirica Karppinen & Krivolutsky 1987S04 [=C. (*Ivarsia) sibiricaS04, CH98]
         `--C. (E.) solhoeyi Colloff 1993 [incl. C. islandica Gjelstrup & Solhoy 1994]S04

Camisia (Camisia) biurus biurus (Koch 1839) [incl. C. exuvialis Grandjean 1939, Nothrus furcatus Koch 1844, Uronothrus kochi Willmann 1943, N. pigerrimus Koch 1844]S04

Camisia (Camisia) biverrucata (Koch 1839) [=Nothrus biverrucatus; incl. C. berlesei Oudemans 1900, C. fischeri Oudemans 1900, C. nicoletii Oudemans 1900]C93

Camisia (Camisia) horrida (Hermann 1804)S04, SC19, S04 [=Notaspis horridaS04, Nothrus horridusBA85, Oribata horridaM98; incl. Nothrus bistriatus Koch 1839S04, Hermannia bistriataS22, Nothrus borealis Thorell 1871S04, Camisia horrida var. borealisS62, Nothris horridus var. borealisT05, Nothrus mutilus Koch 1839S04, Nothrus rugulosus Banks 1895S04, Nothrus runcinatus Koch 1839S04, Nothrus sinuatus Koch 1839S04]

*Camisia (Camisia) segnis (Hermann 1804)S04, SC19, S04 [=Notaspis segnisS04, Nothrus (*Uronothrus) segnisS22, CH98, Oribata segnisCF77; incl. Nothrus bicarinatus Koch 1839S04, Nothrus excisus Banks 1895S04, Nothrus rostratus Koch 1839S04, Camisia segnis segnoides André, Lebrun & Leroy 1984S04, Nothrus ventricosus Koch 1844S04]

Camisia (Camisia) spinifer (Koch 1835)S04 [=Nothrus spiniferS04, N. spiniger (l. c.)CF77; incl. N. ansatus Haupt 1882S04, N. concavus Haupt 1882S04, N. echinatus Koch 1835S04, N. quadricarinatus Haupt 1882S04, N. sordidus Koch 1839S04, N. superbus Berlese 1910S04, Camisia superbaC93, N. taurinus Banks 1906S04]

*Type species of generic name indicated


[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[C93] Colloff, M. J. 1993. A taxonomic revision of the oribatid mite genus Camisia (Acari: Oribatida). Journal of Natural History 27 (6): 1325–1408.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[M98] Michael, A. D. 1898. Oribatidae. In: H. Lohmann (ed.) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen vol. 3. Acarina pp. 1–93. R. Friedländer und Sohn: Berlin.

[SC19] Schaefer, I., & T. Caruso. 2019. Oribatid mites show that soil food web complexity and close aboveground-belowground linkages emerged in the early Paleozoic. Communications Biology 2: 387.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S62] Seyd, E. L. 1962. The moss mites of Kinder Scout, Derbyshire (Acari: Oribatei). Journal of the Linnean Society of London, Zoology 44 (300): 585–591.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

[T05] Trägårdh, I. 1905. Monographie der arktischen Acariden. In: Römer, F., & F. Schaudinn (eds) Fauna Arctica. Eine Zusammenstellun der arktischen Tierformen, mit besonder Berücksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nördliche Eismeer im Jahre 1898 vol. 4 pp. 1–78. Gustav Fischer: Jena.

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