Globularia fluctuata, from here.

Belongs within: Cerithiimorpha.

The Campaniloidea are a group of marine gastropods united by sperm morphology, with members having a euspermatozoon containing seven or eight stright mitochondria, accompanied by a dense segmented sheath. They also share a simple larval shell morphology (Ponder et al. 2008). The Campaniloidea are largely relictual in the modern fauna, with Campanilidae and Ampullinidae each being represented by only a single surviving species, together with the living genus Plesiotrochus.

<==Campaniloidea [Campanilimorpha]
    |--Settsassia Bandel 1992BR05 [SettsassiidaeBR17]
    |    `--*S. obliquecostata (Münster 1841) [=Melania obliquecostata]BR17
    |--Trypanaxis Cossmann 1889BR05 [TrypanaxidaeBR17, Trypanaxinae]
    |    `--*T. umbilicatus (Lamarck 1804) [=Cerithium umbilicatum]BR17
    |--Tylostoma Sharpe 1849BR05 [TylostomatidaeBR17, Tylostomatinae, Tylostominae]
    |    `--*T. torrubiae Sharpe 1849BR17
    |--Vernedia Mazeran 1912 [Vernediidae]BR17
    |    `--*V. laurenti Mazeran 1912BR17
    |    |--ProcampanileTTE93
    |    `--Campanile Bayle 1884 [Campanilinae]BR17
    |         |--*C. leve (Quoy & Gaimard 1834) [=Cerithium leve]BR17
    |         |--C. giganteum [=Cerithium giganteum]W93
    |         `--C. symbolicum Iredale 1917 [incl. C. truncatum Gray 1834]W93
    |    |--Gymnocerithium Cossmann 1906BR17, BR05 [Gymnocerithiidae]
    |    |    `--*G. collegiale (Zittel 1873) [=Cerithium collegiale]S98
    |    `--Diozoptyxis Cossmann 1896 [Diozoptyxidae, Diozoptyxinae, Diozoptyxisinae]BR05
    |         |--*D. monilifera (d’Orbigny 1842) [=Nerinea monilifera]BR17
    |         `--D. marrotiana (d’Orbigny 1842)TTE93
    |    |--Metacerithium Cossmann 1906 [Metacerithiinae, Metacerithinae]BR05
    |    |    |--*M. trimonile (Michelin 1838) [=Cerithium trimonile]BR17
    |    |    `--M. ponsiBR17
    |    `--Nerineopsinae [Nerineopsidae]BR17
    |         |--Nerineopsis Cossmann 1906BR17
    |         |    `--*N. davoustianus (Cotteau 1854) [=Cerithium davoustianum]BR17
    |         `--‘Terebrella’ Andreae 1887 non Maltzan 1886BR17, BR05 [Terebrellidae]
    |              `--*T. guerrei (Hébert & Eudes-Deslongchamps 1860) [=Cerithium guerrei]BR17
    |--Plesiotrochus Fischer 1878BR05 (see below for synonymy)
    |    |--*P. souverbianus Fischer 1878BR17
    |    |--P. acutangulusW93
    |    |--P. crinitus Thiele 1930W93
    |    |--P. fischeri Smith 1909JG19
    |    |--*Hemicerithium’ imperfectumW93
    |    |--‘Tectarius’ luteus Gould 1861R89
    |    |--P. monachus (Crosse & Fischer 1864) [=*Hypotrochus monachus]W93
    |    |--P. pagodiformis Hedley 1907H09
    |    |--P. penetricinctus (Cotton 1932)W93
    |    `--*Trochocerithium’ turritumW93
    |--Gyrodes Conrad 1860BR05 [Gyrodeidae, Gyrodesacea, GyrodidaeBR17, Gyrodinae]
    |    |  i. s.: G. rotundus Stephenson 1941BS00
    |    |--G. (Gyrodes)BR17
    |    |    |--*G. (G.) ‘crenata’ (Conrad 1860) [=Natica crenata non Récluz 1853]BR17
    |    |    `--G. (G.) supraplicata (Conrad 1858)BS00
    |    |--G. (Dockeryella Bandel 1999)BD12
    |    |    |--G. (*D.) major Wade 1926BS00
    |    |    |--G. (D.) euryomphalus (Philippi 1887) [=Natica euryomphalus]BS00
    |    |    |--G. (D.) rennei Kiel & Bandel 2003BD12
    |    |    `--G. (D.) tenellus (Stoliczka 1868)BS00
    |    `--G. (Sohlella)BS00
    |         |--G. (S.) spillmani Gabb 1861BS00
    |         `--G. (S.) yolensis Popenoe, Saul & Suzuki 1987BS00
    `--Ampullinidae [Ampullinoidea, Ampullospiridae]BR17
         |--Cernina fluctuata (Sowerby 1825)TTE93
         |--Amaurellina Bayle in Fischer 1885 [Amaurellinidae]BR05
         |    `--*A. spirata (Lamarck 1804) [=Ampullaria spirata]BR17
         |--Falorina Bandel 2006 [Falorininae]BR17
         |    `--*F. torpediniformis (Böhm 1895) [=Euchrysalis torpediniformis]BR17
         |--Ampullospira Harris 1897 [Ampullospirinae]BR05
         |    |--*A. canaliculata (Morris & Lycett 1854) [=Euspira canaliculata]BR17
         |    `--A. adducta (Phillips 1829)TTE93
         |--Ampullina Bowdich 1822 [Ampullininae]BR05
         |    |--*A. depressa (Lamarck 1804) [=Ampullaria depressa]BR17
         |    `--A. apora (Watson 1881)F27
         |--Globularia Swainson 1840 [Globulariidae, Globulariinae]BR05
         |    |--*G. sigaretina (Lamarck 1804) [=Ampullaria sigaretina, Ampullina sigaretina, Natica sigaretina]BR17
         |    `--G. fluctuata (Sowerby 1825) [incl. Natica imperforata Jay 1836]BC01
         |--Pictavia Cossmann 1925G01
         |    |--*P. pictaviensis (d’Orbigny 1852) [=Natica pictaviensis; incl. N. bajociensis d’Orbigny 1852]G01
         |    |--P. calypso (d’Orbigny 1852) (see below for synonymy)G01
         |    |--P. punctura (Bean 1839) [=Littorina punctura, Natica punctura]G01
         |    `--P. silesiaca (Ahlburg 1906)NE02
         |--Naricopsina Chelot 1886BR05 [=Lobostoma Cossmann 1885 non Gundlach 1840G01; Naricopsinidae, NaricopsininaeBR17]
         |    |--*N. guerangeri (Davoust 1855) [=Neritopsis guerangeri, *Lobostoma guerangeri]BR17
         |    |--N. montreuilensis (Hébert & Eudes-Deslongchamps 1860) (see below for synonymy)G01
         |    |--N. sinuosa Gründel 2001G01
         |    `--N. umbilicata Cossmann 1885G01
         |--Pseudamaura Fischer 1885 [Pseudamauridae, Pseudamaurinae, Pseudamaurinidae]BR05
         |    |--*P. bulbiformis (Sowerby 1832) [=Amaura bulbiformis]BR17
         |    |--P. globosa (Ahlburg 1906)NE02
         |    |--P. gregaria (v. Schlotheim 1822)NE02
         |    |--P. reflecta Etheridge 1902F71
         |    `--P. variablisF71
         `--Oonia Gemmellaro 1878 [incl. Mesospira Cossmann 1892]G01
              |--‘Pseudomelania’ hettangiensis Cossmann 1909 (see below for synonymy)G01
              |--O. acutovata Gründel 2001G01
              |--O. boonei (Cossmann 1924) [=Pseudomelania (Oonia) boonei]G01
              |--O. latiuscula (Morris & Lycett 1851) [=Phasianella latiuscula]G01
              |--O. leymeriei (d’Archiac 1843) [=Phasianella leymeriei, *Mesospira leymeriei]G01
              `--O. subtortilis (Münster 1841)TTE93

Naricopsina montreuilensis (Hébert & Eudes-Deslongchamps 1860) [=Natica montreuilensis; incl. Neritopsis archiaci d’Arch. in Lycett 1863, Natica cornelia Laube 1867, Natica texata Lycett 1863, Lobostoma texatum, Naricopsina texata, L. wastense Cossmann 1885]G01

Pictavia calypso (d’Orbigny 1852) [=Natica calypso, Ampullospira (Pictavia) calypso, N. (Amauropsis) calypso; incl. Pictavia callovica Cossmann 1924]G01

Plesiotrochus Fischer 1878BR05 [incl. Hemicerithium Cossmann 1893W93, Hypotrochus Cotton 1932W93, Trochocerithium Cossmann & Sacco 1896W93; PlesiotrochidaeH96]

‘Pseudomelania’ hettangiensis Cossmann 1909 [=Melania abbreviata Terquem 1855 non Defrance 1823, *Oonia abbreviata]G01

*Type species of generic name indicated


[BD12] Bandel, K., & D. T. Dockery III. 2012. Protoconch characters of Late Cretaceous Latrogastropoda (Neogastropoda and Neomesogastropoda) as an aid in the reconstruction of the phylogeny of the Neogastropoda. Freiberger Forschungshefte C542 (psf 20): 93–128.

[BS00] Bandel, K., & W. Stinnesbeck. 2000. Gastropods of the Quiriquina Formation (Maastrichtian) in central Chile: paleobiogeographic relationships and the description of a few new taxa. Zbl. Geol. Paläont. Teil I 7–8: 757–788.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[G01] Gründel, J. 2001. Neritimorpha und weitere Caenogastropoda (Gastropoda) aus dem Dogger Norddeutschlands und des nordwestlichen Polens. Berliner Geowissenschaftliche Abhandlungen, Reihe E 36: 45–99.

[H96] Healy, J. M. 1996. Molluscan sperm ultrastructure: correlation with taxonomic units within the Gastropoda, Cephalopoda and Bivalvia. In: Taylor, J. D. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 99–113. Oxford University Press: Oxford.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.

[NE02] Nützel, A., & D. H. Erwin. 2002. Battenizyga, a new Early Triassic gastropod genus with a discussion of the caenogastropod evolution at the Permian/Triassic boundary. Paläontologische Zeitschrift 76 (1): 21–27.

[R89] Reid, D. G. 1989. Systematic revision of the Recent species of Peasiella Nevill, 1885 (Gastropoda: Littorinidae), with notes on the fossil species. Nautilus 103 (2): 43–69.

[S98] Stilwell, J. D. 1998. Late Cretaceous Mollusca from the Chatham Islands, New Zealand. Alcheringa 22 (1): 29–85.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[W93] Wilson, B. 1993. Australian Marine Shells vol. 1. Prosobranch Gastropods. Part One. Odyssey Publishing: Kallaroo (Western Australia).

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