Cancellaria

Cancellaria reticulata, copyright Shellnut.

Belongs within: Cancellariidae.

Cancellaria is a genus of relatively large, heavy nutmeg snails with an ovate shell bearing strong columellar plaits.

Narona decaptyx: a fossil vampire
Published 23 March 2018
13.5 mm long specimen of Narona decaptyx, from Landau et al. (2012).

Narona decaptyx was described by Brown & Pilsbry (1911) from a single small, fusiform fossil shell, 11 mm in length, from the Gatun Formation of Panama, north of Panama City. They regarded the formation as probably Oligocene in age but Landau et al. (2012) later referred N. decaptyx to the upper Miocene. Until it’s redescription by the latter, this species was only known from Brown & Pilsbry’s original holotype; Landau et al. described further material from the Bocas del Toro region to the west of the original locality. Both the sites from which N. decaptyx are known are on the Caribbean coast of Panama.

Narona decaptyx is a member of the Cancellariidae, the nutmeg snails. Cancellariids are one of the smaller families of the great neogastropod radiation, the group that also includes such forms as whelks and cone shells. They generally have a more or less developed sculpture of criss-crossing spiral and axial ribs; the latticed pattern this produces is formally referred to as ‘cancellate’ and provides the source of the family’s name. The most distinctive feature of Cancellariidae is their radula, a slender ribbon of long, flexible teeth arranged in a single row. How this radula functioned was long a mystery. Dissections of the gut of cancellariids failed to find any trace of solid food, and it was suggested they may be adapted to some form of suctorial feeding. Some authors suggested that cancellariids might feed by slurping up micro-organisms. Then, in the 1980s, one species of cancellariid Cancellaria cooperi was observed feeding on sleeping electric rays. The snail would cut incisions in the ray’s skin, presumably with its radula, before inserting its proboscis to slurp up the fish’s blood (O’Sullivan et al. 1987). Other cancellariids have been observed feeding on fluids from other invertebrates such as benthic molluscs or their egg masses.

Cancellaria cooperi feeding on an electric ray, copyright Clinton Bauder.

The genus Narona to which N. decaptyx belongs is now restricted to the north Pacific. In this respect, it is not unique. Cancellariids are poorly represented in the modern Caribbean fauna with only six species known from the sea’s shallow waters but they were much more diverse there in N. decaptyx‘s time. However, following the rise of the isthmus of Panama, many cancellariid taxa once found widely in the tropical Americas became extinct for whatever reason on the eastern side of the divide. This happened regularly enough that the term ‘paciphile’ has been coined for referring to such taxa. A number of distinct waves of paciphile extinctions have been identified in the Caribbean cancellariid fossil record and they have been used to identify distinct chronological zones. Narona decaptyx became extinct as part of the GNPMU (Gatunian Neogene Paciphilic Molluscan Unit) 1 period. Other Narona species persisted in the Caribbean and Gulf of Mexico for longer, surviving into the Pliocene, but eventually they too succumbed to whatever dampened this family’s prospects in the region.

Systematics of Cancellaria
Cancellaria Lamarck 1799BR05 (see below for synonymy)
|--*C. (Cancellaria) reticulata (Linnaeus 1767)BR17 (see below for synonymy)
| |--C. r. reticulataPH90
| |--C. r. adelae Pilsbry 1940PH90
| |--C. r. leonensis Mansfield 1930PH90
| `--C. r. leuzingeri Rutsch 1934PH90
|--C. (Euclia Adams & Adams 1854)BC01 [=Hetereuclia (l. c.)PH90, Heteroeuclia Rovereto 1899PH90]
| |--C. (*E.) cassidiformis Sowerby 1832 [=*Heteroeuclia cassidiformis; incl. C. imperialis Michelin 1832]PH90
| |--C. (E.) balboae Pilsbry 1931BC01
| `--C. (E.) laurettae Petit & Harasewych 1998BC01
|--C. (Merica Adams & Adams 1854)W94
| |--C. (*M.) melanostoma Sowerby 1849PH90
| | |--C. m. melanostomaPH90
| | `--C. m. westralis Garrard 1975PH90
| |--C. (M.) asperella Lamarck 1822 [=C. aspercella (l. c.), Merica asprella (l. c.)]PH90
| | |--C. a. asperellaPH90
| | |--‘Merica’ a. okinawana Noda 1980 [=M. asprella (l. c.) okinawana]PH90
| | `--‘Merica’ a. varicosa Shuto 1969PH90
| |--‘Merica’ contorta [incl. M. contorta var. levis Peyrot 1928]PH90
| |--C. (M.) elegans Sowerby 1822W94 (see below for synonymy)
| |--C. (M.) fenestrata Eichwald 1830 (see below for synonymy)PH90
| |--‘Merica’ gerthi Martin 1931PH90
| |--‘Merica’ haweraensis Laws 1940PH90
| |--‘Merica’ kaiparaensis Laws 1939PH90
| |--‘Merica’ kobayashii (Yokoyama 1927) [=Mitra kobayashii]PH90
| |--‘Merica’ promensis Vredenberg 1921PH90
| |--‘Merica’ pukeuriensis Finlay 1930PH90
| |--‘Merica’ salomacensis Peyrot 1928PH90
| `--‘Merica’ variculosa Peyrot 1928PH90
|--C. (Progabbia Dall 1918)PH90
| |--C. (*P.) cooperii (Gabb 1865)ZLK11, PH90
| |--C. (P.) corbicula Dall 1908O27, PH90
| `--C. (P.) io Dall 1896O27
`--C. (Sydaphera Iredale 1929)W94
|--C. (S.) undulata Sowerby 1849W94 (see below for synonymy)
|--C. (S.) anxifer Iredale 1925W94 [=C. purpuriformis anxiferPH90]
|--C. (S.) australis Sowerby 1832W94
|--‘Sydaphera’ delicosa Laseron 1955 [=S. deliciosa (l. c.)]PH90
|--C. (S.) granosa Sowerby 1832W94 [=C. granulosa (l. c.)PH90]
|--‘Sydaphera’ horii Masuda 1967PH90
|--C. (S.) lactea Deshayes 1830 [incl. C. laevigata Sowerby 1832]W94
|--‘Sydaphera’ obnixa Iredale 1936PH90
|--C. (S.) panamuna Garrard 1975W94
`--C. (S.) purpuriformis Kiener 1841 [incl. C. maccoyi Pritchard & Gatliff 1899, C. tasmanica Tenison Woods 1876]W94

Cancellaria incertae sedis:
C. acalypta Woodring 1970PH90
C. acuminata Sowerby 1832PH90
C. acuticarinata Weisbord 1929PH90
C. adicana Pereira Da Costa 1867PH90
C. aegyptiaca Oppenheim 1906PH90
C. aequaeva Ryckholt 1862PH90
C. africana Petit 1970 [=C. imbricata Watson 1882 non Hörnes 1854]PH90
C. agria Manfield 1930PH90
|--C. a. agriaPH90
`--C. a. uaquala Mansfield 1935PH90
C. albida Hinds 1843PH90
C. albrechtina Mayer 1861PH90
C. aldrichi Gardner 1937PH90
C. altavillae Aradas 1846PH90
C. alternata Conrad 1834PH90
C. alternicostula Sacco 1894PH90
C. altispira Gabb 1869PH90
C. alveata Conrad 1833 (see below for synonymy)PH90
C. alveolata Tate 1889PH90
C. ameghinoi von Ihering 1897PH90
C. amoena Olsson & Harbison 1953PH90
C. ampullacera Lesson 1841 (n. d.)PH90
C. anachoreta Mayer 1876PH90
C. ancycla Gardner 1937PH90
C. andersoni Arnold 1909PH90
C. angelana Hanna 1924 [=C. quadrata Moody 1916 non Sowerby 1822]PH90
C. angosturana Marks 1949PH90
C. angulata Eichwald 1830PH90
C. angulifera Deshayes 1864PH90
C. angulovaricosa Sacco 1894PH90
C. angusta Watelet 1851 [=C. angulata (l. c.) nec Eichwald 1830 nec Sieber 1936]PH90
C. annosa Aldrich 1898PH90
C. annulata Stoliczka 1867PH90
C. anomoia Woodring 1970PH90
C. antiqua Dall 1897PH90
C. antiquata Hinds 1843PH90
C. apater Ryckholt 1861PH90
C. aperta Beyrich 1856PH90
C. apimela Woodring 1970PH90
C. aqualica Petit & Harasewych 1986PH90
C. arctica Middendorff 1849PH90
C. arnoldi Dall 1909PH90
C. articularis Sowerby 1832PH90
C. aspera Millet de la Turtaudière 1866PH90
C. atopodonta Petit & Harasewych 1986PH90
C. atraktoides Gardner 1937PH90
C. auriculaperta Vokes 1938PH90
C. auriculoides Millet de la Turtaudière 1854PH90
C. austriaca Hoernes & Auinger 1890PH90
C. babylonica Lea 1833 [incl. C. tera de Gregorio 1890]PH90
C. badrii Abbass 1972PH90
C. bahia Pilsbry & Olsson 1941PH90
C. barjonae Pereira Da Costa 1867 (see below for synonymy)PH90
C. barretti Guppy 1866PH90
C. barystoma Woodring 1970PH90
C. basteroti Deshayes 1864PH90
C. bastropensis Harris 1895PH90
C. baylei Bezançon 1870PH90
C. beata Jung 1965PH90
C. behmi Beyrich 1856PH90
C. bellardii Michelotti 1847PH90
C. beraudiana Millet de la Turtaudière 1866PH90
C. bernardii Mayer 1861 (see below for synonymy)PH90
C. bernayi Cossmann 1889PH90
C. beyrichi Mayer 1859PH90
C. bezanconi de Raincourt 1884PH90
C. bicarinata Hoernes & Auinger 1890PH90
C. bicolor Hinds 1843 [incl. C. septemcostata Odhner 1917]PH90
C. bifoliata Aldrich 1903PH90
C. bifurcoplicata Cossmann 1896PH90
C. binckhorti Nyst 1881 [=C. binkhorsti Cossmann 1899, C. reticulata Binkhorst 1861 non Linné 1767]PH90
C. biplicata Briart & Cornet 1870PH90
C. biplicifera Conrad 1841PH90
C. birchi Addicott 1970PH90
C. birmanica Vredenburg 1921PH90
C. bistriata von Koenen 1889PH90
C. blountiana Mansfield 1935PH90
C. bocageana Crosse & Debeux 1863 [=C. bocagei Dautzenberg & Fischer 1907]PH90
C. bofilli Cossmann 1899 [=C. pulcherrima Almera & Bofill 1898 non Lea 1841]PH90
C. boivini Dareste de la Chavanne 1910PH90
C. boucheti Petit & Harasewych 1986PH90
C. bouczeki Oppenheim 1922 [=Turbinella bouczeki Ržehak 1896 (n. n.)]PH90
C. bournei Hanna 1927PH90
C. boutillieri Cossmann 1889PH90
C. bradleyi Nelson 1870PH90
C. brandenburgi Boettger 1902PH90
C. brauniana Sandberger 1859 [=C. brauneana (l. c.)]PH90
C. brevis Sowerby 1832PH90
C. brocchii Crosse 1861 [incl. C. hirta var. minor Bronn 1848 non C. dufourii var. minor Grateloup 1847]PH90
C. bronnii Bellardi 1841PH90
C. buccinoides Sowerby 1832PH90
C. ‘buccinoides’ von Koenen 1889 nec Sowerby 1832 nec Couthouy 1838PH90
C. bulbulus Sowerby 1832PH90
C. bullata Sowerby 1832 [incl. C. dalli Bartsch 1915]PH90
C. bullbrooki Mansfield 1925 [=C. bulbrooki (l. c.)]PH90
C. burdigalense Peyrot 1928PH90
C. cacellensis Pereira Da Costa 1867PH90
C. calcarata (see below for synonymy)PH90
C. californica Dall 1908PH90
C. callosa Hörnes 1854 (see below for synonymy)PH90
C. caloosahatchiensis Tucker & Wilson 1932PH90
C. calvertensis Martin 1904PH90
C. calvulata Tate 1889PH90
C. calypso White 1887PH90
C. canaliculata Hörnes 1854PH90
C. cancellata (Linné 1758)S11 (see below for synonymy)
|--C. c. cancellataPH90
`--C. c. pluricosticillata Sacco 1894 (see below for synonymy)PH90
C. candida Sowerby 1832PH90
C. caperata Tate 1889PH90
C. capillata Tate 1889PH90
C. carinata Briart & Cornet 1877PH90
C. carolinensis Emmons 1858PH90
C. ‘carolinensis’ Conrad 1863 non Emmons 1858PH90
C. casicalva Marks 1949PH90
C. cassidea (see below for synonymy)PH90
C. centrota Dall 1896PH90
C. cerithea Olsson 1964PH90
C. cerithiopsis Almera & Bofill 1898PH90
C. charapota Olsson 1942PH90
C. charlesworthii Wood 1872PH90
C. chaussyensis Cossmann 1889PH90
C. chinenensis MacNeil 1960PH90
C. cibarcola Anderson 1929PH90
C. cingulata Kaunhowen 1898PH90
C. circumcincta Dall 1873PH90 [=C. (Sveltia) circumcinctaO27, C. circumcarinata (l. c.)PH90]
C. circumspinosa Addicott 1970PH90
C. clathrata Lamarck 1822PH90
C. cloezi Cossmann 1892PH90
C. coaetana Ryckholt 1862PH90
C. coaeva Ryckholt 1862PH90
C. coctilis Reeve 1856PH90
C. codazzii Anderson 1929PH90
C. coensis Mansfield 1930PH90
C. colombiana Olsson 1942PH90
C. cominella Pilsbry & Olsson 1941PH90
C. complicata Handmann 1882PH90
C. condoni Anderson 1905PH90
C. conoidea von Koenen 1885PH90
C. conradiana Dall 1890 [=*Exechoptychia conradiana]PH90
|--C. c. conradianaPH90
`--C. c. clewistonensis Olsson & Harbison 1953PH90
C. contabulata Sowerby 1832 (see below for synonymy)PH90
C. corbula Conrad 1843 (n. d.) [=C. corbulu]PH90
C. coronadosensis Durham 1950PH90
C. coronata Scacchi 1835 [=C. subangulata Scacchi in Dall 1889; incl. C. taeniata Sowerby 1849]PH90
C. corrosa Reeve 1856PH90
C. cossmanni Morlet 1888PH90
C. costulata (Calcara 1840) (see below for synonymy)PH90
C. couvana Vokes 1938PH90
C. crassa Nomland 1917PH90
C. crassicosta Bellardi 1841 (see below for synonymy)PH90
C. crassistria von Koenen 1889PH90
C. crassistriata Wood 1879PH90
C. cremata Hinds 1843PH90
C. crenata Hörnes 1856PH90
C. crenifera Sowerby 1832PH90
C. crenulata Deshayes 1835PH90
C. ‘crenulata’ Adams 1855 non Deshayes 1835PH90
C. crepini Briart & Cornet 1877PH90
C. cretacea Nyst 1881 [=C. obtusa Binkhorst 1861 non Deshayes 1830]PH90
C. crispa Sowerby 1832PH90
C. crossei Semper 1861 [=C. serrata Reeve 1856 non Bronn 1831]PH90
C. crossletensis Covacevich & Frassinetti 1986PH90
C. cruzialis von Ihering 1907PH90
C. cumingiana Petit de la Saussaye 1844 (see below for synonymy)PH90
C. curta von Koenen 1885PH90
C. dalliana Anderson 1905PH90
C. danieli Morlet 1885PH90
C. dariena Toula 1909 (see below for synonymy)PH90
C. darwini Petit 1970BC01
C. dautzenbergi Cossmann 1896PH90
C. decaptyx Brown & Pilsbry 1911PH90
C. ‘decussata’ Bellardi 1841 nec Sowerby 1832 nec Nyst 1838PH90
C. ‘decussata’ Sowerby 1847 nec Sowerby 1832 nec Nyst 1838 nec Bellardi 1841PH90
C. ‘decussata’ Grateloup 1847 nec Sowerby 1832 nec Nyst 1838 nec Bellardi 1841 nec Sowerby 1847PH90
C. defuniak Gardner 1937PH90
C. delecta Deshayes 1864 [=C. elegans Deshayes 1835 non Sowerby 1822]PH90
C. dentifera Deshayes 1864PH90
C. deperdita Michelotti 1861 [incl. C. depertdita var. turrituloides Sacco 1894]PH90
C. depressa Tuomey & Holmes 1856PH90
C. dertocassideum Sacco 1894PH90
C. dertocontorta Sacco 1894 (see below for synonymy)PH90
C. dertonensis Bellardi 1841 (see below for synonymy)PH90
C. dertoscalata Sacco 1894 (see below for synonymy)PH90
C. dertovaricosa Sacco 1894 (see below for synonymy)PH90
C. deshayesana Grateloup 1832 (see below for synonymy)PH90
C. desmotis Gardner 1937PH90
C. desori Mayer 1876PH90
C. deydieri Fontannes 1878PH90
C. diadela Woodring 1970PH90
C. diadema Watelet 1853PH90
C. dinota Woodring 1970PH90
C. distincta Mayer 1876PH90
C. doderleini Mayer 1868 (see below for synonymy)PH90
|--C. d. doderleiniPH90
`--C. d. scabriuscula Moroni 1958PH90
C. dolioides Pilsbry & Olsson 1941PH90
C. doncieuxi Dareste de la Chavanne 1910PH90
C. dregeri Hoernes & Auinger 1890 [incl. C. nassiformis Seguenza 1880 non Lesson 1842]PH90
C. druentica Fontannes 1878PH90
C. druidarum Gardner 1937PH90
C. dubia Deshayes 1864PH90
C. dunkeri Holzapfel 1888PH90
C. duponti Briart & Cornet 1877PH90
C. effosa Handmann 1882PH90
C. egregia von Koenen 1889PH90
C. elata Hinds 1843PH90
C. ellipsis Pilsbry 1922PH90
C. elodiae Carson 1926PH90
C. elongata Nyst 1845PH90
C. elsmerensis English 1914PH90
C. emydis Dall & Ochsner 1928PH90
C. engonata Conrad 1841PH90
C. eogassinense Sacco 1904PH90
C. epidromiformis Tate 1889PH90
C. epistomifera Guppy 1876PH90
|--C. e. epistomiferaPH90
`--C. e. sathra Woodring 1973 [=C. e. lipara Woodring 1970 non C. lipara Woodring 1951]PH90
C. esi Koperberg 1931PH90
C. esmeralda Olsson 1964PH90
C. etheridgei Johnston 1880PH90
C. eucheea Gardner 1947PH90
C. euclethra Maury 1925PH90
C. euetrios Barnard 1959PH90
C. eufaulensis Gabb 1860PH90
C. eutaeniata Cossmann 1896PH90
C. euthymei Barnard 1960PH90
C. exaltata Tate 1889PH90
C. excellens Beyrich 1856 [incl. C. excellens var. rugosior Sacco 1894]PH90
C. exigua Smith 1891PH90
C. exopleura Dall 1908PH90
C. exquisita Preston 1905PH90
C. exumbilicaris Sacco 1894 (see below for synonymy)PH90
C. exwestiana Sacco 1894 (see below for synonymy)PH90
C. fergusoni Carson 1926PH90
C. fernandoensis Arnold 1907 (see below for synonymy)PH90
C. fetzaraensis Dareste de la Chavanne 1910PH90
C. fischeri Adams 1860PH90
C. foveata Almera & Bofill 1884PH90
C. foveolata Sowerby 1849PH90
C. fragosa Olsson 1964PH90
C. frizzelli Marks 1949PH90
C. funerata Conrad 1848PH90
C. funiculata Hinds 1843 (see below for synonymy)PH90
C. funiculifera Vincent in Vincent & Lefèvre 1872PH90
C. fusca Sowerby 1889 (see below for synonymy)PH90
C. fusiformis Cantraine 1835PH90
C. ‘fusiformis’ Deshayes 1864 nec Cantraine 1835 nec Fasciolaria fusiformis Philippi 1845PH90
C. fusospinosa Sacco 1894PH90
C. gabbiana Pilsbry & Johnson 1917PH90
C. gainfarnensis Handmann 1882PH90
C. galei Addicott 1970PH90
C. galvestonensis Harris 1895PH90
C. gaudryi Fontannes 1878PH90
C. gemmata Conrad 1833 (see below for synonymy)PH90
C. gemmulata Sowerby 1832PH90
C. geslini Basterot 1825 (see below for synonymy)PH90
C. gilberti Aldrich 1921PH90
C. glabra Tesch 1915PH90
C. glabricula von Koenen 1894PH90
C. globulosa Holzapfel 1888PH90
C. goniostoma Sowerby 1832 [incl. C. rigida Sowerby 1832]PH90
C. govenderi King 1953PH90
C. gracilenta Wood 1872PH90
C. gracilior Carpenter in Gabb 1869PH90
C. gracilis von Ihering 1897 (see below for synonymy)PH90
C. graciloides Aldrich 1898 [incl. C. graciloides var. bella Aldrich 1898]PH90
C. gradata Hörnes 1854 (see below for synonymy)PH90
C. granifera Deshayes 1830PH90
C. granulata Nyst 1845PH90
C. grateloupi d’Orbigny 1852 [=C. gratteloupi]PH90
C. grayi Tryon 1885PH90
C. grossauensis Handmann 1882PH90
C. guppyi Gabb 1873PH90
C. gurabis Maury 1917PH90
C. haemastoma Sowerby 1832 [=C. haemostoma (l. c.)]PH90
C. hamlini Carson 1926PH90
C. harmeri Glibert 1960PH90
C. harpa von Koenen 1889PH90
C. harpiformis Pilsbry & Olsson 1941PH90
C. harrisi Maury 1917PH90
C. hartti Maury 1925PH90
C. hebertiana Hörnes 1856PH90
C. hemphilli Dall 1909PH90
C. hettneri Anderson 1929PH90
C. hidasensis Hoernes & Auinger 1890PH90
C. hirta (see below for synonymy)PH90
C. hoernesi Kittl 1887PH90
C. hordeola von Koenen 1889PH90
C. hukusimana Nomura & Hatai 1936PH90
C. hypermeces Cossmann 1896PH90
C. igarassuensis Penna 1965PH90
C. imbricata Hörnes 1854 (see below for synonymy)PH90
C. incompta Briart & Cornet 1877PH90
C. inornata Noetling 1901PH90
C. insularis Pilsbry & Johnson 1917PH90
C. intercedens Stoliczka 1867PH90
C. intermedia Bellardi 1841 (see below for synonymy)PH90
C. interrupta Deshayes 1864PH90
C. interstrialis von Koenen 1889PH90
C. irelaniana Cooper 1894PH90
C. islacolonis Maury 1917PH90
C. italica D’Ancona 1872 [incl. C. italica var. crispata Seguenza 1880]PH90
C. iunior Bellardi 1841 [=Sveltia junior]PH90
C. jacksonica Cooke 1926PH90
C. jadisi Olsson 1964PH90
C. japonica Smith 1879PH90
C. jasnini Glibert 1952PH90
C. jayana Keen 1958 [incl. C. clathrata Adams 1855 non Lamarck 1822]PH90
C. jipijapana Pilsbry & Olsson 1941PH90
C. joachimi Handmann 1882PH90
C. joaquinensis Anderson 1905PH90
C. jogjacartensis Martin 1914PH90
C. jonkairiana Nyst 1835 [=C. jonkeiriana (l. c.), C. lajonkairei (l. c.), C. lajonkairii Nyst 1853]PH90
C. jonkeri Koperberg 1931PH90
C. karsteni Anderson 1929PH90
C. keenae Addicott 1970PH90
C. kernensis Addicott 1970PH90
C. kimikoae Hatai 1940PH90
C. kochiensis Katto 1960PH90
C. kugleri Rutsch 1934PH90
C. kunraedensis Kaunhowen 1898PH90
C. labratula von Koenen 1889PH90
C. labrosa Bellardi 1841 (see below for synonymy)PH90
C. lacondamini Olsson 1964PH90
C. lacunosa Hutton 1885PH90
C. laddi Petit 1987 [=C. petiti Ladd 1982 non Olsson 1967]PH90
C. laekeniana Vincent in Cincent & Lefèvre 1872 [=C. laekeniensis (l. c.)]PH90
C. laevescens Guppy 1866 [=C. lavescens (l. c.); incl. C. laevescens var. portoricana Maury 1920]PH90
C. ‘laevigata’ von Koenen 1865 non Sowerby 1832PH90
C. laeviuscula Sowerby 1822PH90
C. lamberti Souverbie in Souverbie & Montrouzier 1870PH90
C. lamyi Koperberg 1931PH90
C. lanceolata Aldrich 1897PH90
C. landesi Van Winkle 1918PH90
C. laqua Mansfield 1935PH90
C. larkinii Nelson 1870PH90
C. latesulcata von Koenen 1885PH90
C. laurensii Grateloup 1832PH90
C. lavelana Hodson in Hodson & Hodson 1931PH90
C. leopoldinae Tournouër in Bouillé 1876PH90
C. lickana Anderson & Martin 1914 [=C. lichana (l. c.)]PH90
C. lima von Koenen 1889PH90
C. limata Yokoyama 1928PH90
C. lipara Woodring 1951PH90
C. lischkei Yokoyama 1926PH90
C. listeri Risso 1826 [=C. lisberi (l. c.)]PH90
C. littoriniformis Sowerby 1832 [=C. littorinaeformis (l. c.)]PH90
C. longispirata Yokoyama 1926PH90
C. losquemadica Maury 1917PH90
C. luscinia Melvill & Standen 1903PH90
C. macnairyensis Sohl 1964PH90
C. macneili Mansfield 1937PH90
C. macrostoma du Bois de Montpéreux 1831 [=C. microstoma (l. c.) non Newton 1895]PH90
C. malachitensis Stanton 1893PH90
C. malaisei Briart & Cornet 1877 [=C. malaisi]PH90
C. maldonadoi Olsson 1964PH90
C. mangelioides Reeve 1856PH90
C. mangyschlakica Bajarunas 1912PH90
C. margaritata Vinassa de Regny 1896PH90
C. marieana Aldrich 1897 [=C. mericana (l. c.)]PH90
C. marksi Olsson 1964PH90
C. martiniana Noetling 1901PH90
C. marylandica Martin 1904PH90
C. matsoni Stephenson 1941PH90
C. mauryae Olsson 1922PH90
C. mediangulata Sacco 1894PH90
C. medinae Philippi 1887PH90
C. menadensis Schepman 1907PH90
C. meridionalis Woods 1906PH90
C. metuloides Olsson 1964PH90
C. michelinii Bellardi 1841 (see below for synonymy)PH90
C. micra Tate 1889PH90
C. microsoma Dall 1908PH90
C. microstoma Newton 1895PH90
C. mihelici Šuklje 1929PH90
C. milletii Deshayes 1830PH90
C. minima Reeve 1856PH90
C. ‘minima’ Geinitz 1874 non Reeve 1856PH90
C. minuta Nyst 1845PH90
C. ‘minuta’ Brown 1849 (n. d.) non Nyst 1845PH90
C. miocaenica Cossmann 1913PH90
C. miocenica Sacco 1894 [=C. miocenicum, C. cassidea var. miocenica Doderlein 1863 (n. n.)]PH90
C. miocrassa Sacco 1894PH90
C. miohirta Sacco 1894PH90
C. miolyrata Sacco 1894PH90
C. mioparva Sacco 1894PH90
C. mioquadrata Sacco 1894PH90
C. mioscalaroides Sacco 1894PH90
C. mississippiensis Conrad 1848PH90
C. mitrodita Gardner 1937PH90
C. mitroides Fischer von Waldheim 1807 (n. d.)PH90
C. modesta Carpenter 1864PH90 [=Cancellaria (Sveltia) modestaO27]
C. modestina Tate 1889PH90
C. moguntina Crosse 1861 [=C. minuta Sandberger 1859 nec Nyst 1845 nec Brown 1849]PH90
C. moharrami Abbass 1972PH90
C. montserratensis Maury 1925PH90
C. moorei Guppy 1866PH90
|--C. m. mooreiPH90
`--C. m. pachia Smith 1940PH90
C. moravica Oppenheim 1922PH90
C. mourloni Briart & Cornet 1877PH90
C. multienensis Morlet 1885 [=C. bezanconi Morlet 1885 non de Raincourt 1885, C. multiensis (l. c.)]PH90
C. multiplicata Lea 1833PH90
C. multistriata Beyrich 1856PH90
C. muratana Nomura & Onisi 1940PH90
C. murayamai Yokoyama 1926 [=C. myrayamai (l. c.)]PH90
C. muricata Risso 1826PH90
C. mutinensis Foresti 1884 (see below for synonymy)PH90
C. mutsuana Hatai, Masuda & Suzuki 1961PH90
C. nana Deshayes 1864PH90
C. nancellaria Woodring 1970PH90
C. nanggulanensis Martin 1914PH90
C. nassariformis Covacevich & Frassinetti 1986PH90
C. nassoides von Koenen 1889PH90
|--C. n. nassoidesPH90
`--C. n. timorensis Koperberg 1931PH90
C. neglecta Michelotti 1861F27b
C. neudorfensis Oppenheim 1922PH90
C. neugeboreni Hörnes 1856PH90
C. nevadensis Anderson & Martin 1914PH90
C. newhallensis Carson 1926PH90
C. nitens Beyrich 1856 [incl. C. nitens var. elatior von Koenen 1889]PH90
C. nitida Adams 1855 [=Scalptia nitida]PH90
C. ‘nitida’ Reeve 1856 non Adams 1855PH90
C. ‘nitida’ von Koenen 1889 nec Adams 1855 nec Reeve 1856PH90
C. notabilis Eichwald 1851 [incl. Pleurotoma cancellatum Eichwald 1830 non Voluta cancellata Linné 1767]PH90
C. nysti Hörnes 1854PH90
C. obesa Sowerby 1832 [incl. C. eburnaeformis Reeve 1856, C. obesa var. planospira Grant & Gale 1931]PH90
C. oblita Michelotti 1861PH90
C. oblonga Sowerby 1825 (see below for synonymy)PH90
C. obsoleta Hörnes 1856 (see below for synonymy)PH90
C. obtusa Deshayes 1830PH90
C. occulta Beyrich 1856PH90
C. ocoyana Addicott 1970PH90
C. oldroydia Carson 1926PH90
C. orbignyana Briart & Cornet 1868PH90
C. oregonensis Conrad 1865 [incl. C. andersoni Clark 1918 non Arnold 1909]PH90
C. ornata Deshayes 1864PH90
C. ‘ornata’ Ovechkin 1954 non Deshayes 1864PH90
C. ‘ornata’ Alekseyev 1963 nec Deshayes 1864 nec Ovechkin 1954]PH90
C. ovata Sowerby 1832PH90
C. ovoidolaevis Sacco 1894PH90
C. ovulum Geinitz 1874PH90
C. pabloensis Clark 1915PH90
C. pacifica Anderson 1905PH90
C. pajana Pilsbry & Olsson 1941PH90
C. pallida Smith 1899PH90
C. palmeri Carson 1926PH90
C. panones Harris 1895 [incl. C. panones var. junipera Harris 1895]PH90
C. papillosa Coppi 1872PH90
C. papyracea Grateloup 1832PH90
C. paraguanensis Hodson in Hodson & Hodson 1931PH90
C. paramoorei Gardner 1937PH90
C. parcestriata Broon in Reiss & Bronn 1862PH90
C. parnensis Cossmann 1896PH90
C. partschi Hörnes 1854PH90
C. parva Lea 1833PH90
C. parvula Beyrich 1856 [incl. C. parvula var. supracosticillata Sacco 1894]PH90
C. paschalis Thiele 1925PH90
C. patricia Thiele 1925PH90
C. patula Mayer 1858PH90
C. patuxentia Martin 1904PH90
C. paucicostata Sowerby 1894PH90
C. pearlensis Meyer & Aldrich in Meyer 1887PH90
C. peninsularis Olsson 1942PH90
C. penita Olsson 1942PH90
C. penrosei Harris 1895 [=C. pemrosei (l. c.)]PH90
C. percostata de Gregorio 1890PH90
C. perdiciana Olsson 1942PH90
C. perrini Carson 1926PH90
C. perspectiva Conrad 1834PH90
|--C. p. perspectivaPH90
`--C. p. alumensis Mansfield 1930PH90
C. petiti Olsson 1967 [=C. cossmanni Olsson 1922 non Morlet 1888]PH90
C. petuchi Harasewych, Petit & Vehecken 1992S11
C. pinguis Gardner 1937PH90
C. pirabensis Maury 1925PH90
C. piscatoria (Gmelin 1791) (see below for synonymy)PH90
C. plagiostoma Conrad 1834PH90
C. planistria von Koenen 1885PH90
C. plebeja Thiele 1925PH90
C. pliocenica Sacco 1894PH90
C. pliopostica Sacco 1894PH90
C. plummeri Olsson 1922PH90
C. plurimixta de Gregorio 1882PH90
C. poetzleinsdorfensis Sieber 1936PH90
C. postypica Sacco 1894PH90
C. posunculensis Anderson & Martin 1914PH90
C. potomacensis Clark & Martin 1901PH90
C. praeevulsa Cossmann 1898 (see below for synonymy)PH90
C. praeindentata Maury 1925PH90
C. praetiosa Oppenheim 1922PH90
C. priama Harris 1895PH90
C. pristinaPH90
|--C. p. pristinaPH90
`--C. p. acutiplicata Shuto 1962PH90
C. producta Sowerby 1903PH90
C. propegemmata de Gregorio 1890PH90
C. propevenusta Mansfield 1929PH90
C. prophylactica Icke & Martin 1907PH90
C. propinqua Kaunhowen 1898PH90
C. protrigonostoma Sacco 1894PH90
C. proxima Risso 1826 (n. d.)PH90
C. prunicola Martin 1904PH90
C. pseudocancellata Noetling 1901PH90
C. pseudoevulsa d’Orbigny 1852PH90
C. pulcherrima Lea 1841PH90
C. pulchra Sowerby 1932PH90
C. puruensis Martin 1914PH90
C. puschi Semper 1861 (see below for synonymy)PH90
C. ‘puschi’ Hoernes & Auinger 1890 non Semper 1861PH90
C. pycta Olsson 1964PH90
C. pygmaea Adams 1852PH90
C. pyrgota Edwards in Lowry 1866PH90
C. quasilla Petit 1987 [=C. cretacea Smith 1899 non Nyst 1881]PH90
C. quercollis [incl. C. quercollis var. greggi Harris 1899]PH90
C. ramonensis Clark 1918PH90
C. rapa Nomland 1917PH90
C. rapella Johnson 1904PH90
C. rara Aoki 1954PH90
C. raulini Mayer 1858PH90
C. reticulatoides Martin 1904PH90
C. reyesi Covacevich & Frassinetti 1986PH90
C. rhabdota Bayan 1873 [=C. canaliculata Deshayes 1864 non Hörnes 1854]PH90
C. rhombea von Koenen 1889PH90
C. richardpetiti Petuch 1987PH90
C. ricinus Pecchioli 1864PH90
C. ringens Sandberger 1859 [=C. cingens (l. c.)]PH90
C. rothi Semper 1861 [=C. nodulifera Beyrich 1856 non Sowerby 1825]PH90
C. rotunda Dall 1892 [=C. conradiana var. rotunda]PH90
|--C. r. rotundaPH90
`--C. r. floridana Olsson & Petit 1964PH90
C. rowelli Dall in Guppy & Dall 1896PH90
C. rudis Whitfield 1892PH90
C. rudolphi Covacevich & Frassinetti 1986PH90
C. rugosa Lamarck 1822PH90
C. ‘rugosa’ von Koenen 1889 non Lamarck 1822PH90
C. runchaena Gardner 1937PH90
C. saccoi Hoernes & Auinger 1890PH90
C. sanctaemariae Carson 1926PH90
C. sandbergeri Tournouër 1879PH90
C. santiagensis Marks 1949PH90
C. scabra Deshayes 1830 (see below for synonymy)PH90
C. scalariformis Lamarck 1822PH90
C. scalarina Lamarck 1822PH90
C. scalaroides Wood 1857PH90
C. scalata Sowerby 1832 [incl. C. tenuis Adams 1855]PH90
C. scalatella Guppy 1873PH90
C. scheibei Anderson 1929PH90
C. schroeckingeri Hoernes & Auinger 1890PH90
C. schucherti Olsson 1932PH90
C. schythei Philippi 1855PH90
C. scobina Hedley & Petterd 1906PH90 [=Admete scobinaF27a]
C. scrobiculata Hörnes 1854 [incl. C. scrobiculatum var. costicillatina Sacco 1894]PH90
C. semiclathrata Morlet 1885PH90
C. semicostata Tate 1889PH90
C. semidisjuncta Sowerby 1849PH90
C. semipellucida Adams & Reeve 1850PH90
C. semota Jung 1969PH90
C. semperi Speyer 1867PH90
C. semperiana Crosse 1863PH90
C. sendoi Hatai 1941PH90
C. separata Deshayes 1864PH90
C. septemlirata Gabb 1860PH90
C. sericea Dall 1892PH90
C. serramata Olsson 1967 [=C. serrata Olsson 1964 nec Bronn 1831 nec Reeve 1856]PH90
C. serrata Bronn 1831 (see below for synonymy)PH90
C. shilohensis Pilsbry & Harbison 1933PH90
C. shirleyae Olsson 1967PH90
C. siletzensis Anderson in Hanna 1924 (see below for synonymy)PH90
C. silvestris Wrigley 1935PH90
C. simiana Hanna 1924 [=C. crassa Waring 1917 non Nomland 1917]PH90
C. ‘similis’ Aradas 1846 non Sowerby 1833PH90
C. ‘similis’ Giebel 1847 nec Sowerby 1833 nec Aradas 1846PH90
C. simplex Anderson 1905PH90
C. simulata von Koenen 1889PH90
C. sinuosa Cossmann 1889PH90
C. siouxensis Erickson 1974PH90
C. sismondai D’Ancona 1872PH90
C. smithvillensis Harris 1895PH90
C. smocki Weller 1907PH90
C. sobrantensis Clark 1918PH90
C. soriensis Eames 1952PH90
C. sotoensis Aldrich 1908PH90
C. souverbiei Crosse 1868PH90
C. spatiosa Nelson 1870PH90
C. speciosa Deshayes 1864PH90
C. spectabilis Deshayes 1864PH90
C. specularis Watson 1882PH90
C. spellenbergi Riedel 1932PH90
C. spengleriana Deshayes 1830 [incl. C. tritonis Sowerby 1832]PH90
|--C. s. spenglerianaPH90
`--C. s. koreanica Hatai & Kotaka 1952PH90
C. sphaericula Cossmann 1889PH90
C. sphenoidostoma Gardner 1937PH90
C. spherotopleura Gardner 1937PH90
C. spinifera Grateloup 1832 [=C. spirifer (l. c.)]PH90
C. spinosa Grateloup 1827PH90
C. springvaleensis Mansfield 1925PH90
C. stantoni Dickerson 1913PH90
C. stibara Gardner 1937PH90
C. striatulata Deshayes 1835 [=C. striatula (l. c.)]PH90
C. strombecki Speyer 1864PH90
C. stromboides Grateloup 1832PH90
C. strongi Shasky 1961PH90
C. strozzii Pecchioli 1864PH90
C. stuardoi McLean & Andrade 1982PH90
C. subacuminata d’Orbigny 1852 (see below for synonymy)PH90
C. subalta Conrad 1869PH90
C. subangulosa Wood 1848 [incl. C. subangulosa var. rotundata von Koenen 1889]PH90
C. subcancellata d’Orbigny 1852PH90
C. subcarinata Bronn 1831PH90
C. subcylindrica von Koenen 1889PH90
C. subevulsa d’Orbigny 1850PH90
C. subhirta d’Orbigny 1852 [=C. battersbyi Bell 1870]PH90
C. subsuturalis d’Orbigny 1852 [=C. suturalis Grateloup 1832 non Sowerby 1822]PH90
C. subthomasiae Dall 1890PH90
C. subtiarophora Gardner 1937PH90
C. subtilicancellata Maury 1925PH90
C. subumbilicata Grateloup 1847 [=C. varicosa var. subumbilicata, C. subvaricosa d’Orbigny 1852]PH90
C. succineiformis Boettger 1906PH90
C. suessi Hoernes 1875PH90
C. sulcata Bellardi 1841PH90
C. supercretacea Favre 1869PH90
C. suppar Ryckholt 1862PH90
C. surpacifica Olsson 1967 [=C. pacifica Pilsbry & Olsson 1941 non Anderson 1905]PH90
C. sursalta Marks 1949PH90
C. suturalis Sowerby 1822 [=C. biplex Sowerby 1822; incl. C. ranelliformis Chenu 1859]PH90
C. sylvaerupis Harris 1897 [=C. silvaerupis]PH90
C. synchrona Ryckholt 1862PH90
C. tabatai Yokoyama 1926PH90
C. tabulata Gardner & Aldrich 1919PH90
C. taiwanensis Nomura 1935PH90
C. tapeina Woodring 1970PH90
C. taurangulifera Sacco 1894PH90
C. taurinia Bellardi 1841 (see below for synonymy)PH90
C. tauroconvexula Sacco 1894 (see below for synonymy)PH90
C. taurocrassum Sacco 1894 [=C. taurapertum Sacco 1894; incl. C. taurocrassum var. subauriculata Sacco 1894]PH90
C. taurofoveolata Sacco 1894 [=C. taurofaveolata]PH90
C. taurolaevigatum Sacco 1894 (see below for synonymy)PH90
C. taurospinulosa Sacco 1894PH90
C. telemba Olsson 1964PH90
C. ‘tenuis’ Reeve 1856 non Adams 1855PH90
C. tenuistriata von Koenen 1865PH90
C. terebralis von Koenen 1889PH90
C. terraereginensis Etheridge 1907PH90
C. terryi Olsson 1942PH90
C. teschi Koperberg 1931PH90
C. testaventricosa Grateloup 1847PH90
C. textilis Kiener 1841PH90
C. thiemeana Geinitz 1874PH90
C. tholoensis Ladd 1934PH90
C. thomasiana Crosse 1861 [=C. thomasiae (l. c.); incl. C. scalarina Sowerby 1849 non Lamarck 1822]PH90
C. thuringiae Giebel 1853PH90
C. tjibaliungensis Martin 1895PH90
C. toroensis Olsson 1922PH90
C. torquayensis Chapman 1922PH90
C. torquilla Zekeli 1852PH90
C. tortiplica Conrad 1865 [incl. C. tortiplica var. subevulsopsis de Gregorio 1890]PH90
C. tortoniana Sacco 1894PH90
C. torula Weisbord 1962PH90
C. trapezium Borson 1820PH90
C. trema Olsson 1932PH90
C. triangularis Nelson 1870PH90
C. tricincta von Koenen 1885 [=Admete bicincta (l. c.)]PH90
C. trigonostomaH43
C. trilineata Handmann 1882PH90
C. tritonidea Gabb 1866PH90
C. trochilia Olsson 1964PH90
C. trochlearis Faujas de Saint Fond 1817 (see below for synonymy)PH90
C. tuberculosa Sowerby 1832 (see below for synonymy)PH90
C. tumescens von Koenen 1889PH90
C. tumida von Koenen 1889PH90
C. turneri White 1889PH90
C. turriculata Tate 1889PH90
C. ‘turrita’ Sowerby 1874 non C. nodulosa var. turrita Bellardi 1841PH90
C. turritissima Meyer 1886PH90
C. tyosenensis Hatai & Nisiyama 1940PH90
C. ulmula Harris 1895PH90
C. umbilicaris (see below for synonymy)PH90
C. umbilicata Lesson 1842 (n. d.)PH90
C. uniangulata Deshayes 1830 (see below for synonymy)PH90
C. urceolata Hinds 1843PH90
C. urcianensis d’Ancona 1872PH90
C. urumacoensis Hodson in Hodson & Hodson 1931PH90
C. varicifera Tenison-Woods 1879PH90
C. varicosissima Handmann 1882PH90
C. venezuelana Hodson in Hodson & Hodson 1931PH90
C. ventricosa Hinds 1843 [incl. C. affinis Adams 1852]PH90
C. venusta Tuomey & Holmes 1856PH90
C. verbeeki Martin 1895PH90
C. verreauxii Kiener 1841 [=C. verauxii (l. c.), C. verrauxii (l. c.)]PH90
C. vespertina Anderson 1905PH90
C. vetusta Gabb 1866PH90
C. vidali Philippi 1887PH90
C. vindobonensis Sacco 1894PH90
C. waltoniana Gardner 1937PH90
C. wannonensis Tate 1889PH90
C. washingtonensis Weaver 1916PH90
C. waynensis Mansfield 1940PH90
C. weaveri Etherington 1931PH90
C. werenfelsi Jung 1965PH90
C. westziana Grateloup 1847 [=C. westiana (l. c.); incl. C. westiana var. tauroparvula Sacco 1894]PH90
C. wigginsi Emerson & Hertlein 1964PH90
C. wilmeri Sowerby 1881PH90
C. wynoochensis Weaver 1916 [=C. wynoocheensis (l. c.), C. wynootchensis]PH90
C. xenia Olsson 1964PH90
C. yolandia Pilsbry & Olsson 1941PH90
C. yonabaruensis MacNeil 1960PH90
C. zahni Böse 1910PH90
C. zapoteca Böse 1910PH90
C. zitteli Wanner 1902PH90

Nomina nuda: Cancellaria abnormis Gray 1847PH90
Cancellaria alvaniopsis Harris in Cossmann 1899PH90
Cancellaria antiqua Wagner in Bronn 1848 non Dall 1897PH90
Cancellaria aspera Edwards in Newton 1891 non Millet de la Turtaudière 1866PH90
Cancellaria aveniformis Edwards in Newton 1891PH90
Cancellaria azumai Habe in Azuma 1960PH90
Cancellaria berolinensis Beyrich 1847PH90
Cancellaria bonacorsi Doderlein in Davoli 1982PH90
Cancellaria cassiniana Newton 1891PH90
Cancellaria ciliata Michaelis & Scherk 1847PH90
Cancellaria clarendonensis Edwards in Newton 1891PH90
Cancellaria concinna Wood 1842PH90
Cancellaria continua Tate in Tate & Dennant 1893PH90
Cancellaria cornigera Braun 1851PH90
Cancellaria dewalquei Nyst in Dewalque 1868PH90
Cancellaria dictyella Cossmann 1899PH90
Cancellaria dubia Edwards in Wrigley 1935 non Deshayes 1864PH90
Cancellaria echinata Doderlein in Sacco 1894PH90
Cancellaria eximia Edwards in Newton 1891PH90
Cancellaria fontinalis Tate in Tate & Dennant 1893PH90
Cancellaria glabrata Bosquet in Vincent 1886PH90
Cancellaria globularis Edwards in Jones 1878PH90
Cancellaria gracilenta Edwards in Newton 1891 non Wood 1872PH90
Cancellaria kobayasii Otuka 1937PH90
Cancellaria latilabris Bon. in Michelotti 1838PH90
Cancellaria lauta Tate in Tate & Dennant 1893PH90
Cancellaria laviae Hoffmann in Brugnone 1880PH90
Cancellaria macrostoma Doderlein 1863 non du Bois de Montpéreux 1831PH90
Cancellaria scabriuscula var. major Pallary 1900 nec C. dufourii var. major Grateloup 1847 nec C. gracilis var. major von Ihering 1899PH90
Cancellaria marysvillensis Dickerson 1916PH90
Cancellaria hirta var. minor Almera & Bofill 1892 nec C. dufourii var. minor Grateloup 1847 nec C. hirta var. minor Bronn 1848 nec C. evulsa var. minor Beyrich 1856 nec C. cancellata var. minor Almera & Bofill 1884 nec C. fusca var. minor Sowerby 1889 nec C. piscatoria var. minor Pallary 1920PH90
Cancellaria scabriuscula var. minor Pallary 1900 nec C. dufourii var. minor Grateloup 1847 nec C. hirta var. minor Bronn 1848 nec C. evulsa var. minor Beyrich 1856 nec C. cancellata var. minor Almera & Bofill 1884 nec C. fusca var. minor Sowerby 1889 nec C. piscatoria var. minor Pallary 1920PH90
Cancellaria ampullacea var. multicostata Bellardi 1841 nec C. uniangulata var. multicostata Bellardi 1841 nec C. nodulosa var. multicostata Bellardi 1841 nec C. buccinula var. multicostata Bellardi 1841PH90
Cancellaria multilinea Edwards in Newton 1891PH90
Cancellaria muricata Wood 1847 non Risso 1826PH90
Cancellaria mutica Basterot in Grateloup 1832PH90
Cancellaria namnetica Vasseur 1881PH90
Cancellaria nana Doderlein in Davoli 1980 non Deshayes 1864PH90
Cancellaria nassaeformis Wood in Newton 1891PH90
Cancellaria nodigera Edwards in Newton 1891PH90
Cancellaria ozawai Otuka 1937PH90
Cancellaria pinolensis Clark 1918PH90
Cancellaria polygona Bellardi 1840PH90
Cancellaria porrecta Edwards in Newton 1891PH90
Cancellaria pseudaustralis Tate in Dennant 1889PH90
Cancellaria pulchra Briart & Cornet in Dewalque 1868 non Sowerby 1832PH90
Cancellaria pupa Edwards in Newton 1891PH90
Cancellaria raulini Grateloup in Peyrot 1928 non Mayer 1858PH90
Cancellaria reticulata Edwards in Newton 1891 nec Voluta reticulata Linné 1767 nec C. reticulata Binkhorst 1861 [=C. eoreticosa Cossmann 1899]PH90
Cancellaria rosea Beck in Sowerby 1849PH90
Cancellaria roydonensis Edwards in Newton 1891PH90
Cancellaria scalaris Michelotti 1838PH90
Cancellaria sismondiana Grateloup in Peyrot 1928PH90
Cancellaria spirata Edwards in Newton 1891 non Lamarck 1822PH90
Cancellaria spirulata Doderlein in Davoli 1982PH90
Cancellaria staringi Bosquet in Staring 1860PH90
Cancellaria subglobosa Mighels 1841J49
Cancellaria sublaevis Bellardi 1840PH90
Cancellaria tenuiplica Edwards in Newton 1891PH90
Cancellaria turris Edwards in Newton 1891PH90
Cancellaria turritella Hoeninghaus 1831PH90
Cancellaria typica Krause in Patel 1888PH90
Cancellaria unifasciata Orbigny in Dall 1909PH90
Cancellaria venatrix Edwards in Newton 1891PH90
Cancellaria westi Bast. in Mayer 1872PH90
Cancellaria wildi Mayer 1872PH90
Cancellaria ytenensis Edwards in Newton 1891PH90

Cancellaria Lamarck 1799BR05 [=Cancellarius Montfort 1810PH90, Plicaria Fabricius 1823 (nom. inv.) non Fuckel 1870 (ICBN)PH90; incl. Exechoptychia Cossmann 1903PH90]

Cancellaria alveata Conrad 1833 [incl. C. leai Crosse 1861, C. tessellata Lea 1833 non Sowerby 1832, C. sculptura Lea 1833, C. sculpturata (l. c.)]PH90

Cancellaria barjonae Pereira Da Costa 1867 [incl. C. barjonae var. bearnensis Peyrot 1928, C. barjonae var. dertoturrita Sacco 1894, C. doderleini Moroni 1958 non Mayer 1868, C. barjonae var. laevior Sacco 1894, C. barjonae var. megastoma Peyrot 1928, C. barjonae var. perhirta Sacco 1894, C. barjonae var. salomacensis Peyrot 1928, C. barjonae var. spiratior Sacco 1894, C. barjonae var. subaspinosa Peyrot 1928, C. barjonae var. tauroaspinosa Sacco 1894]PH90

Cancellaria bernardii Mayer 1861 [incl. C. bernardii var. tauroaspira Sacco 1894, C. bernardii var. taurocaudata Sacco 1894]PH90

Cancellaria calcarata [incl. C. calcarata var. ampleumbilicata Sacco 1894, C. calcarata var. basicosticillata Sacco 1894, C. calcarata var. dertopercostata Sacco 1894, C. calcarata var. quadrulata Almera & Bofill 1884, C. calcarata var. scalaratula Sacco 1894, C. calcarata var. spinosissima Sacco 1894, C. calcarata var. tauroconnectens Sacco 1894]PH90

Cancellaria callosa Hörnes 1854 [incl. C. callosa var. cacellotransiens Sacco 1894, C. callosa var. taurolaevior Sacco 1894, C. callosa var. tauroturrita Sacco 1894]PH90

Cancellaria cancellata (Linné 1758)S11 [=Voluta cancellataPH90, Murex scabriculus Linnaeus 1758 non Voluta scabriculus Linnaeus 1758PH90, M. scabriusculus (l. c.)PH90; incl. Cancellaria cancellata var. astensis Bellardi 1841PH90, Canc. costata Sowerby 1822PH90, Canc. cancellata var. dertonensis Bellardi 1841 non C. bonellii var. dertonensis Bellardi 1841PH90, Canc. cancellata var. infracosticillata Sacco 1894PH90, Canc. cancellata var. minor Almera & Bofill 1884 nec C. dufourii var. minor Grateloup 1847 nec C. hirta var. minor Bronn 1848 nec C. evulsa var. minor Beyrich 1856PH90, Canc. cancellata var. perscalarata Sacco 1894PH90, Canc. cancellata var. praecedens Beyrich 1856PH90, Canc. cancellata var. pseudonassoides Sacco 1894PH90, Buccinum pyrozonias Gmelin 1791PH90, Buccinella rotundata Perry 1811PH90, Cancellaria cancellata var. suprafasciata Sacco 1894PH90, Canc. cancellata var. taurinia Bellardi 1841 nec C. varricosa var. taurinia Bellardi 1841 nec C. lyrata var. taurinia Bellardi 1841 nec C. uniangulata var. taurinia Bellardi 1841 nec C. nodulosa var. taurinia Bellardi 1841 nec C. bonelli var. taurinia Bellardi 1841 nec C. evulsa var. taurinia Bellardi 1841PH90, Canc. cancellata var. tauroelongata Sacco 1894PH90, Cantharus triplicatus Röding 1798PH90]

Cancellaria cancellata pluricosticillata Sacco 1894 [incl. C. cancellata pleuricosticillata anom. costicillatissima Sacco 1894 non C. tribulus var. costicillatissima Sacco 1894, C. cancellata pluricosticillata subvar. elatocostata Sacco 1894, C. cancellata pluricosticillata subvar. elatocosticillata Sacco 1894, C. cancellata pleuricosticillata anom. irregularisuturata Sacco 1894, C. cancellata pluricosticillata anom. variecosticillata Sacco 1894]PH90

Cancellaria cassidea [=C. expidea Tate 1889 (n. n.); incl. C. cassidea var. excassidea Sacco 1894, C. cassidea var. paucicostata Sacco 1894 non C. paucicostata Sowerby 1894, C. cassidea var. spiratior Sacco 1894]PH90

Cancellaria contabulata Sowerby 1832 [incl. C. forestieri Montrouzier in Souverbie & Montrouzier 1863, C. montrouzieri Souverbie in Souverbie & Montrouzier 1863, C. pusilla Sowerby 1832, C. rougeyroni Souverbie in Souverbie & Montrouzier 1870]PH90

Cancellaria costulata (Calcara 1840) [=Buccinum costulatus non C. costulata Lamarck 1803 (not preoc. if in dif. gen.), C. costata (l. c.) nec Sowerby 1822 nec Sowerby 1833 nec Lea 1833]PH90

Cancellaria crassicosta Bellardi 1841 [incl. C. crassicosta var. gracilina Sacco 1894, C. crassicosta var. percostatula Sacco 1894]PH90

Cancellaria cumingiana Petit de la Saussaye 1844 [=Trigonostoma cumingii (l. c.); incl. C. cumingiana var. subobtusa Crosse 1863]PH90

Cancellaria dariena Toula 1909 [=C. darienensis Cossmann 1913; incl. C. dariena var. trachyostraca Brown & Pilsbry 1911]PH90

Cancellaria dertocontorta Sacco 1894 [incl. C. dertocontorta var. eximbricata Sacco 1894, C. dertocontorta var. rotundulina Sacco 1894]PH90

Cancellaria dertonensis Bellardi 1841 [=C. bonellii var. dertonensis; incl. C. dertonensis var. angulatina Sacco 1894, C. dertonensis var. compressospira Sacco 1894, C. dertonensis var. dertonassoides Sacco 1894, C. dertonensis var. subbiplicata Sacco 1894, C. dertonensis var. tauroacosticillata Sacco 1894, C. dertonensis var. taurobliquata Sacco 1894, C. dertonensis var. taurocompressa Sacco 1894, C. dertonensis var. tauroturris Sacco 1894]PH90

Cancellaria dertoscalata Sacco 1894 [incl. C. dertoscalata var. exscrobiculata Sacco 1894, C. dertoscalata var. hirtocostata (l. c.), C. dertoscalata var. pseudotribulus Sacco 1894, C. dertoscalata var. suprafasciolata Sacco 1894, C. dertoscalata var. tricostata Sacco 1894]PH90

Cancellaria dertovaricosa Sacco 1894 [inc. C. dertovaricosa var. magnoturrita Sacco 1894, C. dertovaricosa var. spinulatior Sacco 1894, C. dertovaricosa var. subasuturata Sacco 1894]PH90

Cancellaria deshayesana Grateloup 1832 [=C. deshayesi (l. c.), C. deshayesiana (l. c.); incl. C. deshayesiana (l. c.) var. longonassoides Sacco 1894, C. deshayesiana (l. c.) var. mericoides Sacco 1894, C. deshayesiana (l. c.) var. tauropercostata Sacco 1894 non C. ampullacea var. tauropercostata Sacco 1894]PH90

Cancellaria doderleini Mayer 1868 [incl. C. doderleini var. convexior Sacco 1894, C. doderleini var. laticostata Sacco 1894 non C. laticostata Tenison-Woods 1880, C. doderleini var. raricosta Sacco 1894, C. doderleini var. turritula Sacco 1894]PH90

Cancellaria exumbilicaris Sacco 1894 [=C. umbilicaris var. exumbilicaris, Trigonostoma bellardii de Stefani & Pantanelli 1878 non Cancellaria bellardii Michelotti 1847]PH90

Cancellaria exwestiana Sacco 1894 [incl. C. exwestiana var. compressa Sacco 1894, C. exwestiana var. fusoascalaris Sacco 1894, C. exwestiana var. perproducta Sacco 1894, C. exwestiana var. pyramidata Sacco 1894, C. exwestiana var. tuberculosa Sacco 1894 non C. tuberculosa Sowerby 1832]PH90

Cancellaria fernandoensis Arnold 1907 [incl. C. fernandoensis var. tribulis Nomland 1917, C. tribulus (l. c.) non Voluta tribulus Brocchi 1814]PH90

Cancellaria funiculata Hinds 1843 [=C. fumiculata (l. c.), C. punctulata (l. c.); incl. C. lyrata Adams & Reeve 1850 non Voluta lyrata Brocci 1814]PH90

Cancellaria fusca Sowerby 1889 [incl. C. fusca var. minor Sowerby 1889 nec C. dufourii var. minor Grateloup 1847 nec C. hirta var. minor Bronn 1848 nec C. evulsa var. minor Beyrich 1856 nec C. cancellata var. minor Almera & Bofill 1884]PH90

Cancellaria gemmata Conrad 1833 [incl. C. costata Lea 1833 nec Sowerby 1822 nec Sowerby 1833, C. impressa Conrad 1865, C. lirata Conrad 1865, C. plicata Lea 1833]PH90

Cancellaria geslini Basterot 1825 [incl. C. geslini var. compressohirta Sacco 1894, C. geslini var. crassonana Sacco 1894, C. geslini var. crassopostica Sacco 1894, C. geslini var. elongata Grateloup 1847 non C. elongata Nyst 1845, C. geslini var. exgeslini Sacco 1894, C. geslini var. ornatissima Boettger 1906, C. geslini var. taurofasciata Sacco 1894, C. geslini var. taurotransiens Sacco 1894, C. geslini var. ventricosa Grateloup 1847 non C. ventricosa Hinds 1843]PH90

Cancellaria gracilis von Ihering 1897 [incl. C. gracilis var. major von Ihering 1899 non C. dufourii var. major Grateloup 1847, C. gracilis var. observatoria von Ihering 1907]PH90

Cancellaria gradata Hörnes 1854 [incl. C. gradata var. dertumbilicata Sacco 1894, C. gradata var. exampullacea Sacco 1894, C. gradata var. masferreri Almera & Bofill 1884, C. gradata var. tauroconnectens Sacco 1894 non C. calcarata var. tauroconnectens Sacco 1894]PH90

Cancellaria hirta [incl. C. hirta var. colligens Sacco 1894 non C. inermis var. colligens Sacco 1894, C. hirta var. dertosimplex Sacco 1894, C. nodulosa var. maior Bellardi 1841, C. hirta var. major nec C. dufourii var. major Grateloup 1847 nec C. gracilis var. major von Ihering 1899, C. hirta var. obsoleta Brugnone 1880 non C. obsoleta Hörnes 1856, C. hirta var. parva Brugnone 1880 nec C. parva Lea 1833 nec Philippi 1860, C. hirta var. peracuminata Sacco 1894, C. hirta var. suboblitecostata Sacco 1894]PH90

Cancellaria imbricata Hörnes 1854 [incl. C. imbricatum var. crassocostata Sacco 1894, C. imbricata var. dertocosticillata Sacco 1894]PH90

Cancellaria intermedia Bellardi 1841 [=C. nodulosa var. intermedia Bellardi 1840 (n. n.); incl. C. intermedia var. afasciata Sacco 1894, C. intermedia var. plioligustica Sacco 1894]PH90

Cancellaria labrosa Bellardi 1841 [=Fusus lebrosus Bellardi 1839 (n. n.); incl. C. labrosa var. altavillae Libassi 1859 non C. altavillae Aradas 1846, C. altavillae var. strictoturrita Sacco 1894]PH90

Cancellaria michelinii Bellardi 1841 [incl. C. michelinii var. compressula Sacco 1894 non C. serrata var. compressula Sacco 1894, C. michelinii var. costatissima Sacco 1894, C. michelinii var. errata Sacco 1894, C. michelinii var. ovatoventricosa Sacco 1894, C. michelinii var. percostatoacuta Sacco 1894, C. michelinii var. percrassulata Sacco 1894]PH90

Cancellaria mutinensis Foresti 1884 [incl. C. mutinensis var. cacellowestiana Sacco 1894, C. mutinensis var. dertinflata Sacco 1894, C. mutinensis var. pyramidatospira Sacco 1894]PH90

Cancellaria oblonga Sowerby 1825 [=C. elongata (l. c.) nec Nyst 1845 nec C. geslini var. elongata Grateloup 1847; incl. C. bifasciata Deshayes 1830, C. decussata Nyst 1838 non Sowerby 1832]PH90

Cancellaria obsoleta Hörnes 1856 [incl. C. obsoleta var. taurocostatior Sacco 1894, C. obsoleta var. taurolatior Sacco 1894]PH90

Cancellaria piscatoria (Gmelin 1791) [incl. C. piscatoria var. crassicosticillata Sacco 1894, C. nodulosa var. laevilabris Bellardi 1841, C. piscatoria var. minor Pallary 1920 nec C. dufourii var. minor Grateloup 1847 nec C. hirta var. minor Bronn 1848 nec C. evulsa var. minor Beyrich 1856 nec C. cancellata var. minor Almera & Bofill 1884 nec C. fusca var. minor Sowerby 1889, C. nodulosa var. multicostata Bellardi 1841 non C. uniangulata var. multicostata Bellardi 1841, C. nodulosa Lamarck 1822, C. piscatoria var. pseudotypica Sacco 1894, Murex semilunaris Gmelin 1791, C. nodulosa var. taurinia Bellardi 1841 nec C. varricosa var. taurinia Bellardi 1841 nec C. lyrata var. taurinia Bellardi 1841 nec C. uniangulata var. taurinia Bellardi 1841, C. piscatoria var. tranhirta (l. c.), C. piscatoria var. transhirta Sacco 1894, C. nodulosa var. turrita Bellardi 1841]PH90

Cancellaria praeevulsa Cossmann 1898 [=Bonellitia praeevulsa Cossmann 1899, C. similis Kaunhowen 1898 nec Sowerby 1833 nec Aradas 1846 nec Giebel 1847]PH90

Cancellaria puschi Semper 1861 [incl. Voluta citharella Brongniart 1823 non Cancellaria citharella Lamarck 1822, C. cytharella (l. c.)]PH90

Cancellaria scabra Deshayes 1830 [incl. C. scabra var. dertoparva Sacco 1894, C. scabra var. taurocosticillata Sacco 1894]PH90

Cancellaria serrata Bronn 1831 [incl. C. serrata var. aequicosticillata Sacco 1894, C. serrata var. compressula Sacco 1894, C. serrata var. pauciserrata Sacco 1894, C. serrata var. simplicicostata Sacco 1894, C. serrata f. squarrosa Davoli 1980]PH90

Cancellaria siletzensis Anderson in Hanna 1924 [=C. rotunda Anderson & Martin 1914 non C. conradiana var. rotunda Dall 1892]PH90

Cancellaria subacuminata d’Orbigny 1852 [=C. acuminata Bellardi 1841 non Sowerby 1832; incl. C. subacuminata var. convexior Sacco 1894 non C. doderleini var. convexior Sacco 1894, C. subacuminatum var. costulatior Sacco 1894]PH90

Cancellaria taurinia Bellardi 1841 [=C. varricosa var. taurinia; incl. C. taurinia var. exvaricosa Sacco 1894, C. taurinia var. mioconnectens Sacco 1894, C. taurinia var. multistriata Bellardi 1840 (n. n.) non C. multistriata Beyrich 1856, C. taurinia var. parvoturrita Sacco 1894]PH90

Cancellaria tauroconvexula Sacco 1894 [incl. C. tauroconvexula var. infracosticillata Sacco 1894 non C. cancellata var. infracosticillata Sacco 1894]PH90

Cancellaria taurolaevigatum Sacco 1894 [incl. C. taurolaevigatum var. cancellatina Sacco 1894 non C. evulsa var. cancellatina Sacco 1894, C. taurolaevigatum var. laevior Sacco 1894 non C. barjonae var. laevior Sacco 1894]PH90

Cancellaria trochlearis Faujas de Saint Fond 1817 [incl. C. trochlearis var. disparstriata Hölzl 1958, C. trochlearis var. gravecostata Hölzl 1958, C. trochlearis var. intermedia Hölzl 1958 non C. intermedia Bellardi 1841, C. trochlearis var. pseudovilia Sacco 1894]PH90

Cancellaria tuberculosa Sowerby 1832 [=C. tuberculata (l. c.); incl. C. lobata Swainson 1840, C. trinodosa Chenu 1859]PH90

Cancellaria umbilicaris [incl. C. umbilicaris var. ligustica Sacco 1894, C. umbilicaris var. parvotriangula Sacco 1894, C. umbilicaris var. spinosella Sacco 1894, C. umbilicaris var. subcanaliculata Grateloup 1847]PH90

Cancellaria uniangulata Deshayes 1830 [incl. C. uniangulata var. bicosticillata Sacco 1894 non C. varicosa var. bicosticillata Sacco 1894, C. uniangulata var. dertolyratoides Sacco 1894, C. uniangulata var. dertopercostulata Sacco 1894, C. fusulus Bronn 1831, C. fusus (l. c.), C. uniangulata var. multicostata Bellardi 1841, C. uniangulata var. pluricostulata Sacco 1894, C. uniangulata var. pseudocoronata Sacco 1894, C. uniangulata var. taurinia Bellardi 1841 nec C. varricosa var. taurinia Bellardi 1841 nec C. lyrata var. taurinia Bellardi 1841]PH90

*Cancellaria (Cancellaria) reticulata (Linnaeus 1767)BR17 [=Voluta reticulataBR17, *Cancellarius reticulatusPH90, *Plicaria reticulataPH90; incl. Cancellaria reticulata var. alba Webb 1938 (n. n.)PH90, Buccinella canulata Perry 1811PH90, Voluta reticulata var. reeveiP79]

Cancellaria (Merica) elegans Sowerby 1822W94 [=C. reeveana Crosse 1861PH90, C. reevana (l. c.)PH90; incl. C. reeveana var. laticosta Löbbecke 1881PH90, C. latecostata (l. c.)PH90, C. laticostata (l. c.) nec Tenison-Woods 1880 nec C. doderleini var. laticostata Sacco 1894PH90, C. loebbeckei Kuroda & Habe 1971PH90, C. reeveana var. subsinensis Löbbecke 1881PH90]

Cancellaria (Merica) fenestrata Eichwald 1830 [incl. C. fenestrata var. depressocostata Sacco 1894, C. fenestrata var. dertopostica Sacco 1894, Merica fenestrata var. gracilis Friedberg 1914, M. fenestrata var. paucicostata Friedberg 1914, M. fenestrata var. rotundata Friedberg 1914, Cancellaria fenestrata var. supracostulata Sacco 1894, C. fenestrata var. tauracuta Sacco 1894, C. fenestrata var. tauroelongata Sacco 1894 non C. cancellata var. tauroelongata Sacco 1894]PH90

Cancellaria (Sydaphera) undulata Sowerby 1849W94 [incl. Sydaphera delicosa Laseron 1955PH90, W94, S. deliciosa (l. c.)PH90, S. obnixa Iredale 1936PH90, W94, *S. renovata Iredale 1929PH90, C. undulata var. truncata Sowerby 1849PH90]

*Type species of generic name indicated

References

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

Brown, A. P., & H. A. Pilsbry. 1911. Fauna of the Gatun Formation, Isthmus of Panama. Proceedings of the Academy of Natural Sciences of Philadelphia 63 (2): 336–373.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[H43] Hinds, R. B. 1843. On new species of Cancellaria. Proceedings of the Zoological Society of London 11: 47–49.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213–231.

Landau, B., R. E. Petit & C. M. da Silva. 2012. The family Cancellariidae (Mollusca: Gastropoda) in the Neogene of the Bocas del Toro region, Panama, with the description of seven new species. Journal of Paleontology 86 (2): 311–339.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 1. Stanford University Press: Stanford University (California).

O’Sullivan, J. B., R. R. McConnaughey & M. E. Huber. 1987. A blood-sucking snail: the Cooper’s nutmeg, Cancellaria cooperi Gabb, parasitizes the California electric ray, Torpedo californica Ayres. Biological Bulletin 172 (3): 362–366.

[PH90] Petit, R. E., & M. G. Harasewych. 1990. Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia). Nautilus 103 (Suppl. 1): 1–69.

[P79] Petterd, W. F. 1879. List of the Australian volutes. Journal of Conchology 2: 340–345.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

[W94] Wilson, B. 1994. Australian Marine Shells vol. 2. Prosobranch Gastropods. Part Two (Neogastropods). Odyssey Publishing: Kallaroo (Western Australia).

[ZLK11] Zou, S., Q. Li & L. Kong. 2011. Additional gene data and increased sampling give new insights into the phylogenetic relationships of Neogastropoda, within the caenogastropod phylogenetic framework. Molecular Phylogenetics and Evolution 61: 425–435.

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