Cardium costatum, copyright Joop Trausel and Frans Slieker.

Belongs within: Cardiinae.

Cardium is a genus of cockles with radial ribs strongly expressed internally and spines modified into hollow keels (Schneider 2002).

<==Cardium Linnaeus 1758L58
    |--C. aculeatumH04
    |--C. bechei Reeve 1847H09
    |--C. biangulatumC64
    |--C. biradiatum Bruguière 1789H09
    |--C. blandum Gould 1850 [=C. (Cerastoderma) blandum]S02
    |--C. californiense Deshayes 1839 [=C. (Cerastoderma) californiense]S02
    |--C. cardissa Linnaeus 1758L58
    |--C. carolinensis Conrad 1863 [=C. (Cerastoderma) carolinensis]S02
    |--C. chipolanum Dall 1900 [=C. (Cerastoderma) chipolanum]S02
    |--C. ciliare Linnaeus 1758L58
    |--C. ciliatum Fabricius 1780 [=C. (Cerastoderma) ciliatum]S02
    |--C. citrinumG79 [incl. C. serratum Linnaeus 1758G79, L58]
    |--C. consorsC64
    |--C. corallinum Linnaeus 1758L58
    |--C. corbis [incl. C. californianum Conr. 1837]C64
    |--C. costatum Linnaeus 1758S02
    |--C. craticuloides Conrad 1845 [=C. (Cerastoderma) craticuloides]S02
    |--C. cygnorumM54
    |--C. decoratum Grewingk 1850 [=C. (Cerastoderma) decoratum]S02
    |--C. deshayesiPP64
    |--C. dianthinum Melvill & Standen 1899H09
    |--C. druidicum Dall 1900 [=C. (Cerastoderma) druidicum]S02
    |--C. dupuchense Reeve 1845H09
    |--C. eburniferumG79
    |--C. edule Linnaeus 1758L58 [incl. C. rusticum Eichwald 1841 non Linnaeus 1758S01]
    |--C. elatumC64
    |--C. elongatum Bruguière 1789H09
    |--C. exiguum [incl. C. pygmaeum]J64b
    |--C. fasciatumN79
    |--C. fittoniG76
    |--C. fornicatum Sowerby 1841H09
    |--C. fragum Linnaeus 1758H09
    |--C. gabbii Remond 1863C64
    |--C. glaucumG84
    |--C. graniferumC64
    |--C. haitenseG79
    |--C. humanum Linnaeus 1758L58
    |--C. hungerfordiP98
    |--C. imbricatum Sowerby 1841H09
    |--C. impressum Lightfoot 1786K65
    |--C. indicumC64
    |--C. isocardia Linaneus 1758L58
    |--C. lacunosum Reeve 1845H09
    |--C. laevigatum Linnaeus 1758H09
    |--C. lamarckiiCS77
    |--C. laperoussiiC64
    |--C. leptopleura Conrad 1845 [=C. (Cerastoderma) leptopleura]S02
    |--C. leucostomum [incl. C. marmoreum, C. subelongatum]G79
    |--C. linteum Conrad 1855C64
    |--C. lobulatum Deshayes 1855H09
    |--C. lyratum Sowerby 1841H09
    |--C. maculatumC64
    |--C. maculosum Wood 1815H09
    |--C. magnum Linnaeus 1758L58
    |--C. medium Linnaeus 1758L58
    |--C. minimum [incl. C. loveni, C. suecicum]J64a
    |--C. modestum Conrad 1855 [=Fulvia modesta]C64
    |--C. multispinosum Sowerby 1840H09
    |--C. multistriatum Sowerby 1833S02
    |--C. muricatum Linnaeus 1758L58
    |--C. muticumBRW98
    |--C. nicolletiC64
    |--C. nodosumN79
    |--C. norvegicumM62
    |    |--C. n. norvegicumM62
    |    `--C. n. senegalensisM62
    |--C. nuttallii Conrad 1837 [=C. (Cerastoderma) nuttallii]S02
    |--C. obovaleC64
    |--C. oviputamenG79
    |--C. pansatrum Dall 1900 [=C. (Cerastoderma) pansatrum]S02
    |--C. papillosumM62
    |--C. paucicostatumPP64
    |--C. pectinatum Linnaeus 1758L58
    |--C. phlyctaena Dall 1900 [=C. (Cerastoderma) phlyctaena]S02
    |--C. procerumC64
    |--C. productum Deshayes 1855H09
    |--C. protrusum Lightfoot 1786K65
    |--C. pseudofossile Reeve 1844 [=C. (Cerastoderma) pseudofossile]S02
    |--C. rackettiM54
    |--C. reevianum Dunker 1852H09
    |--C. rubicundum Reeve 1844H09
    |--C. rusticum Linnaeus 1758L58
    |--C. senticosumC64
    |--C. serricostatum Melvill & Standen 1899H09
    |--C. (Protocardium) serumH86
    |--C. solidum Linnaeus 1758L58
    |--C. spatiosumH86
    |--C. spinosum Lightfoot 1786K65
    |--C. stultorum Linnaeus 1758L58
    |--C. taphrium Dall 1900 [=C. (Cerastoderma) taphrium]S02
    |--C. transcendens Melvill & Standen 1899H09
    |--C. triste Linnaeus 1758L58
    |--C. tuberculatum Linnaeus 1758L58
    |--C. unedo Linnaeus 1758H09
    |--C. variegatum Sowerby 1841H09
    |--C. virgineum Linnaeus 1758L58
    `--C. waltonianum Dall 1900 [=C. (Cerastoderma) waltonianum]S02

*Type species of generic name indicated


[BRW98] Beesley, P. L., G. J. B. Ross & A. Wells (eds) 1998. Fauna of Australia vol. 5. Mollusca: The Southern Synthesis. Part A. Australian Biological Resources Study: Canberra.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[CS77] Cramp, S., & K. E. L. Simmons (eds) 1977. Handbook of the Birds of Europe, the Middle East and North Africa: The Birds of the Western Palaearctic vol. 1. Ostrich to Ducks. Oxford University Press: Oxford.

[G64] Gosling, E. M. 1984. The systematic status of Mytilus galloprovincialis in western Europe: a review. Malacologia 25 (2): 551–568.

[G76] Grigorescu, D. 1976. Paratethyan seals. Systematic Zoology 25 (4): 407–419.

[G79] Guppy, R. J. L. 1879. First sketch of a marine invertebrate fauna of the Gulf of Paria and its neighbourhood. Part I.—Mollusca. Journal of Conchology 2: 151–172.

[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H86] Hutton, F. W. 1886. The Mollusca of the Pareora and Oamaru systems of New Zealand. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 205–237.

[J64a] Jeffreys, J. G. 1864a. Report of the committee appointed for exploring the coasts of Shetland by means of the dredge. Report of the British Association for the Advancement of Science 33: 70–81.

[J64b] Jeffreys, J. G. 1864b. The Upper Tertiary fossils at Uddevalla, in Sweden. Report of the British Association for the Advancement of Science 33 (Notices and Abstracts): 73–79.

[K65] Kay, E. A. 1965. The Reverend John Lightfoot, Daniel Solander, and the Portland Catalogue. Nautilus 79 (1): 10–19.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[M54] Macpherson, J. H. 1954. Molluscs (sea shells and snails). In: Willis, J. M. (ed.) The Archipelago of the Recherche pp. 55–63. Australian Geographical Society: Melbourne.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[N79] Norman, A. M. 1879. The Mollusca of the fiords near Bergen, Norway. Journal of Conchology 2: 8–77.

[PP64] Peres, J. M., & J. Picard. 1964. Nouveau manuel de bionomie benthique de la mer Mediterranee. Recueil des Travaux de la Station Marine d’Endoume, Bulletin 31 (27): 5–137.

[P98] Prothero, D. R. 1998. Bringing Fossils to Life: An introduction to paleobiology. WCB McGraw-Hill: Boston.

[S02] Schneider, J. A. 2002. Phylogeny of cardiid bivalves (cockles and giant clams): revision of the Cardiinae and the importance of fossils in explaining disjunct biogeographical distributions. Zoological Journal of the Linnean Society 136: 321–369.

[S01] Sturany, R. 1901. Csigák és kagylók [Mollusken]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 69–74. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

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