Variegated limpets
Cellana tramoserica, copyright John Turnbull.

Belongs within: Patellogastropoda.

Cellana is a tropical Indo-Pacific genus of limpets with a solid, strongly ribbed shell (Knight et al. 1960).

Characters (from Knight et al. 1960): Shell fairly solid, apex subcentral; radial ribs strong; interior brilliantly glazed.

<==Cellana Adams 1869 P61 [incl. Bertinia Jousseaume 1883 BR05, Helcioniscus Dall 1871 KC60; Bertinidae, Bertiniidae]
|–C. capensis (Gmelin 1791) NO04
`–+–+–C. toreuma (Reeve 1854) NO04
|  `–+–C. pricei Powell 1973 NO04
|     `–C. taitensis (Röding 1798) NO04
`–+–+–C. orientalis (Pilsbry 1891) NO04
|  `–+–C. grata (Gould 1859) NO04
|     `–C. nigrolineata (Reeve 1854) NO04 [incl. *Bertinia bertinia Jousseaume 1883 BR05]
`–+–C. testudinaria (Linnaeus 1758) NO04
`–+–C. solida (Blainville 1825) NO04
`–C. tramoserica (Holten 1802) NO04

Cellana incertae sedis:
*C. cernica (Adams 1869) [=Nacella cernica] KC60
C. ampla Lindberg & Hickman 1986 TTE93
C. denticulata (Martyn 1784) P61
C. ornata (Dillwyn 1817) P61
C. radians (Gmelin 1791) P61 [=Helcioniscus radians F27; incl. Patella decora F27]
|–C. r. radians P61
|–C. r. flava (Hutton 1873) P61
`–C. r. perana Iredlae 1915 P61
C. radiata (Born 1778) BD86
C. stellifera (Gmelin 1791) P61
|–C. s. stellifera P61
`–C. s. phymatius (Suter 1905) P61
C. strigilis (Hombron & Jacquinot 1841) P61
|–C. s. strigilis P61
|–C. s. bollonsi Powell 1955 P61
|–C. s. chathamensis (Pilsbry 1891) P61
|–C. s. flemingi Powell 1955 P61
|–C. s. oliveri Powell 1955 P61
`–C. s. redimiculum (Reeve 1854) P61
‘Helcioniscus’ variegatus H15

*Type species of generic name indicated


[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117–141.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[H15] Hedley, C. 1915. Presidential address. Journal and Proceedings of the Royal Society of New South Wales 49 (1): 1–77, pls 1–7.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[NO04] Nakano, T., & T. Ozawa. 2004. Phylogeny and historical biogeography of limpets of the order Patellogastropoda based on mitochondrial DNA sequences. Journal of Molluscan Studies 70: 31–41.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

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