Celtidaceae

Chinese hackberry Celtis sinensis, copyright Geographer.

Belongs within: Rosales.

The Celtidaceae, hackberries, are shrubs or trees with unisexual (sometimes bisexual) flowers, drupaceous fruits, a curved embryo, and the cotyledons mostly variously folded (Soepadmo 1977, as Celtideae).

<==Celtidaceae [Celteae, Celtidoideae]
    |--Aphananthe Planch. 1848 (nom. cons.) (see below for synonymy)S77
    |    |--A. aspera [=Homoioceltis aspera]S77
    |    |--A. cuspidata (Bl.) Planch. in DC. 1873 (see below for synonymy)S77
    |    `--A. philippinensis Planch. 1848 (see below for synonymy)S77
    |--Gironniera Gaudich. 1844 [incl. Helminthospermum Thw. 1854, Nemostigma Planch. 1848]S77
    |    |--G. celtidifolia Gaudich. 1844 (see below for synonymy)S77
    |    |--G. hirta Ridl. 1920S77
    |    |--G. nervosa Planch. 1848 [incl. G. penangiana Gandog. 1919, G. sponioides Gandog. 1919]S77
    |    |--G. parvifolia Planch. 1848 (see below for synonymy)S77
    |    |--G. pyriformisJD05
    |    |--G. rhamnifolia Bl. 1856S77
    |    `--G. subaequalis Planch. 1848 (see below for synonymy)S77
    |--Trema Lour. 1790S77
    |    |--T. angustifolia (Planch.) Bl. 1856 (see below for synonymy)S77
    |    |--T. aspera [incl. Sponia virgata]S77
    |    |--T. cannabina Lour. 1790 (see below for synonymy)S77
    |    |--T. guineensisS77
    |    |--T. lamarckianaJ87
    |    |--T. micranthaJ87
    |    |--T. orientalis (Linné) Bl. 1856 (see below for synonymy)S77
    |    |--T. politoriaS77
    |    `--T. tomentosa (Roxb.) Hara 1971 (see below for synonymy)S77
    |         |--T. t. var. tomentosaLK14
    |         `--T. t. var. asperaLK14
    |--Ampelocera ruiziiS77
    |--Chaetacme aristidaS77, BP60
    |--HemipteleaS77
    |--LozanellaS77
    |--Parasponia Miq. 1851S77
    |    |--P. andersonii (Planch.) Planch. in DC. 1873 (see below for synonymy)S77
    |    |--P. melastomatifolia Smith 1914 [incl. P. simulans Merr. & Perry 1941]S77
    |    |--P. parviflora Miq. 1851 [incl. P. similis Bl. 1856]S77
    |    |--P. rigida Merr. & Perry 1941S77
    |    `--P. rugosa Bl. 1856 [incl. P. aspera Bl. 1856, Trema philippinensis Elmer 1934]S77
    |--PteroceltisS77
    |--Zelkova serrataS77, LO98 [incl. Zelkowa keakiLO98]
    `--Celtis Linné 1753 [incl. Solenostigma Endl. 1833]S77
         |--C. australisR-CT01 [incl. C. tournefortiiH91]
         |    |--C. a. var. australisS77
         |    `--C. a. var. eriocarpaS77
         |--C. begonioidesHL08
         |--C. hildebrandii Soepadmo 1977S77
         |--C. iguaneaGJ09
         |--C. laevigataS77
         |--C. latifolia (Bl.) Planch. in DC. 1873 (see below for synonymy)S77
         |--C. luzonica Warb. 1905S77
         |--C. occidentalisWM09
         |--C. pallidaPB83
         |--C. paniculata (Endl.) Planch. 1848 (see below for synonymy)S77
         |--C. philippensis Blanco 1837 (see below for synonymy)S77
         |    |--C. p. var. philippensisS77
         |    `--C. p. var. wightii (Planch.) Soepadmo 1977 (see below for synonymy)S77
         |--C. reticulataH93
         |--C. rigescens (Miq.) Planch. 1873 (see below for synonymy)S77
         |--C. rubrovenia Elmer 1908 [incl. C. similis Merr. & Perry 1941]S77
         |--C. sinensisHQC03
         |    |--C. s. var. sinensisLO98
         |    `--C. s. var. japonicaLO98
         |--C. spinosaS77
         |--C. talaS06
         |--C. tetrandra Roxb. 1832 (see below for synonymy)S77
         |--C. timorensis Span. 1841 (see below for synonymy)S77
         `--C. tupalangiS77

Aphananthe Planch. 1848 (nom. cons.) non Link 1821 [incl. Galumpita Bl. 1856, Gironniera subg. Galumpita Hook. f. 1880, Homoioceltis Bl. 1856, Mirandaceltis Sharp 1958]S77

Aphananthe cuspidata (Bl.) Planch. in DC. 1873 [=Cyclostemon cuspidatum Bl. 1825, Galumpita cuspidata (Bl.) Bl. 1856, Gironniera cuspidata (Bl.) Kurz 1877; incl. Gi. curranii Merr. 1909, Aphananthe lissophylla Gagnep. 1925, Gi. lucida Kurz 1877, Gi. nitida Bth. 1861, Gi. reticulata Thw. 1861, Gi. thompsoni King ex Cowan & Cowan 1929, Gi. yunnanensis Hu 1940, Aphananthe yunnanensis (Hu) Grudz. 1964]S77

Aphananthe philippinensis Planch. 1848 [incl. Taxotrophis rectinervia F.v.M. 1863, A. rectinervia (F.v.M.) Planch. in DC 1873]S77

Celtis latifolia (Bl.) Planch. in DC. 1873 [=Solenostigma latifolium Bl. 1856; incl. C. kajewskii Mer.. & Perry 1941, S. zippelii Bl. 1856, C. zippelii (Bl.) Planch. in DC. 1873]S77

Celtis paniculata (Endl.) Planch. 1848 [=Solenostigma paniculatum Endl. 1833; incl. C. ingens F.v.M. 1864, C. pacifica Planch. 1848, C. vitiensis Smith 1943]S77

Celtis philippensis Blanco 1837 [=C. philippinensis (l. c.), Solenostigma philippinensis (Blanco) Miq. 1859; incl. So. brevinerve Bl. 1856, Celtis brevinervis (Bl.) Planch. in DC. 1873, C. collinsae Craib. 1918, Solenostigma consimile Bl. 1856, C. philippensis var. consimile (Bl.) Leroy 1948, So. laurifolium var. constricta Miq. 1859, So. djungiel Bl. 1856, Celtis djungiel (Bl.) Planch. in DC. 1873, So. hasseltii Bl. 1856, C. hasseltii (Bl.) Planch. in DC. 1873, So. laurifolium Bl. 1856, C. laurifolia (Bl.) Planch. in DC. 1873, C. mauritiana Planch. 1848, C. mindanaensis Elmer 1915, C. multifolia Elmer 1939 (nom. inv.), Sponia strychnifolia Teysm. & Binn. 1853, C. strychnoides Planch. 1848]S77

Celtis philippensis var. wightii (Planch.) Soepadmo 1977 [=C. wightii Planch. 1848, Solenostigma wightii (Planch.) Miq. 1859]S77

Celtis rigescens (Miq.) Planch. 1873 [=Solenostigma rigescens Miq. 1861; incl. C. asperifolia Merr. 1920, C. nymanii K.Sch. in K.Sch. & Laut. 1905, Solenostigma sumatrana Miq. 1861, C. sumatrana (Miq.) Planch. in DC. 1873]S77

Celtis tetrandra Roxb. 1832 [=Sponia tetrandra (Roxb.) Voigt 1845, incl. C. acata Hamilt. 1834, C. glabra Planch. 1848, C. hamiltonii Planch. 1848, C. tetrandra var. hamiltonii Hook. f. 1888, C. mollis Planch. 1848, C. tetrandra var. mollis (Planch.) Hook. f. 1888, C. napalensis Planch. 1848, C. roxburghii Planch. 1848, C. serotina Planch. 1848, C. trinervia Roxb. 1832 non Lamk. 1797]S77

Celtis timorensis Span. 1841 [incl. C. cinnamomea Lindl. ex Planch. 1848, C. crenato-serrata Merr. 1910, C. dysodoxylon Thw. 1861, C. hamata Bl. 1856, C. reticulosa Miq. 1851, C. waitzii Bl. 1856]S77

Gironniera celtidifolia Gaudich. 1844 [incl. G. grandifolia Merr. & Perry 1939, G. retinervia Merr. & Perry 1939, G. sibuyanensis Elmer 1913]S77

Gironniera parvifolia Planch. 1848 [=G. parvifolium; incl. G. subaequalis var. ceylanica Planch. 1848, G. paucinervia Merr. 1917, Helminthospermum scabridum Thw. in Hook. 1854, G. scabrida (Thw.) Alston in Trimen 1931 non G. subaequalis var. scabrida Bl. 1856, G. zeylanica Gandog. 1919]S77

Gironniera subaequalis Planch. 1848 [=G. nervosa var. subaequalis (Planch.) Kurz 1877; incl. G. amboinensis Laut. 1913, Sponia annulata Teysm. & Binn. 1855, G. blumei Gandog. 1919, G. borneensis Gandog. 1919, G. subaequalis var. brevistylis Bl. 1856, G. chinensis Bth. 1861, G. costata Miq. 1855, G. ferruginea Gandog. 191, G. longifolia Craib 1918, G. subaequalis var. papuana Smith 1912, G. subaequalis var. scabrida Bl. 1856, G. subaequalis var. serrulata Bl. 1856, G. sumatrana Gandog. 1919]S77

Parasponia andersonii (Planch.) Planch. in DC. 1873 [=Sponia andersonii Planch. 1848; incl. P. paucinervia Merr. & Perry 1939]S77

Trema angustifolia (Planch.) Bl. 1856 [=Sponia angustifolia Planch. 1848; incl. S. acuminatissima Miq. 1861, Trema acuminatissima (Miq.) Boerl. 1900, T. orientalis var. bicolor De Wit 1949, T. lanceolata Merr. 1931, Sponia sampsonii Hance 1866]S77

Trema cannabina Lour. 1790 [incl. Celtis amboinensis Willd. 1805, Sponia amboinensis (Willd.) Decne 1834, Trema amboinensis (Willd.) Bl. 1856, T. orientalis var. amboinensis (Willd.) Kurz 1877, T. carinata Bl. 1856, Sponia carinata (Bl.) Miq. 1859, T. timorensis var. carinata (Bl.) Laut. 1913, Celtis commersonii Brongn. in Duperrey 1829, Sponia commersonii (Brongn.) Decne 1834, Trema commersonii (Brongn.) Bl. 1856, S. glabrescens Planch. 1848, T. glabrescens (Planch.) Bl. 1856, S. virgata var. major Planch. 1848, Trema morifolia Bl. 1856, S. morifolia (Bl.) Planch. in DC. 1873, T. pallida Bl. 1856, S. pallida (Bl.) Miq. 1859, T. timorensis var. pallida (Bl.) Laut. 1913, T. pubigera Bl. 1856, Sponia pubigera (Bl.) Miq. 1859, T. virgata var. pubigera (Bl.) Laut. 1913, T. cannabina var. scabra, T. virgata var. scabra, Sponia timorensis Decne 1834, T. timorensis (Decne) Bl. 1856, S. vieillardii Planch. in DC. 1873, T. vieillardii (Planch.) Schltr 1905, S. virgata Planch. 1848, T. virgata (Planch.) Bl. 1856, S. viridis Planch. 1848, S. aspera var. viridis (Planch.) Bth. 1873, Trema viridis (Planch.) Bl. 1856]S77

Trema orientalis (Linné) Bl. 1856 [=Celtis orientalis Linné 1753, Sponia orientalis (L.) Decne 1834; incl. S. argentea Planch. 1848, Trema argentea (Planch.) Bl. 1865, T. orientalis var. bicornis De Wit 1949, T. burmannii Bl. 1865, Sponia burmannii (Bl.) Planch. in DC. 1873, Celtis discolor Brongn. in Duperry 1829, S. discolor (Brongn.) Decne 1834, Trema discolor (Brongn.) Bl. 1865, Celtis rigida Bl. 1825, Sponia rigida (Bl.) Decne 1834, Trema orientalis var. rigida (Bl.) Laut. 1913, T. rigida (Bl.) Bl. 1865, T. scaberrima Bl. 1865, Sponia scaberrima (Bl.) Miq. 1859, S. wightii Planch. 1848, Trema wightii (Planch.) Bl. 1865]S77

Trema tomentosa (Roxb.) Hara 1971 [=Celtis tomentosa Roxb. 1832, Sponia tomentosa (Roxb.) Planch. 1848; incl. S. blancoi Planch. 1848, Trema blancoi (Planch.) Bl. 1856, T. dielsiana Hand.-Mazz. 1929, Sponia griffithii Planch. 1848, T. griffithii (Planch.) Bl. 1856, T. imbricata Bl. 1856, Sponia imbricata (Bl.) Planch. in DC. 1873, S. velutina Planch. 1848, Trema velutina (Planch.) Bl. 1856]S77

*Type species of generic name indicated

References

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[GJ09] Gagné, R. J., & M. Jaschhof. 2009. Cecidomyiidae (gall midges). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 293–314. NRC Research Press: Ottawa.

[HQC03] Hadiah, J. T., C. J. Quinn & B. J. Conn. 2003. Phylogeny of Elatostema (Urticaceae) using chloroplast DNA data. Telopea 10 (1): 235–246.

[HL08] Henniger, M., & R. Leder. 2008. Eozäne Makroflora der Querfurter Mulde. Mauritiana 20 (2): 229–251.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[H91] Hubálek, Z. 1991. Biogeographic indication of natural foci of tick-borne infections. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 255–260. SPB Academic Publishing: The Hague.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.

[JD05] Junge, F. W., M. Dolezych, H. Walther, T. Böttger, A. Kühl, L. Kunzmann, P. Morgenstern, T. Steinberg & R. Stange. 2005. Ein Fenster in Landschaft und Vegetation vor 37 Millionen Jahren: Lithologische, sedimentgeochemische und paläobotanische Befunde aus einem Paläoflusssystem des Weißelsterbeckens. Mauritiana 19 (2): 185–273.

[LO98] Lack, H. W., & H. Ohba. 1998. Die Xylothek des Chikusai Kato. Willdenowia 28: 263–276.

[LK14] Lyons, M. N., G. J. Keighery, L. A. Gibson & T. Handasyde. 2014. Flora and vegetation communities of selected islands off the Kimberley coast of Western Australia. Records of the Western Australian Museum Supplement 81: 205–244.

[PB83] Price, M. V., & J. H. Brown. 1983. Patterns of morphology and resource use in North American desert rodent communities. Great Basin Naturalist Memoirs 7: 117–134.

[R-CT01] Ragusa-di Chiara, S., & H. Tsolakis. 2001. Phytoseiid faunas of natural and agricultural ecosystems in Sicily. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 522–529. CSIRO Publishing: Melbourne.

[S77] Soepadmo, E. 1977. Ulmaceae. Flora Malesiana, Series I—Spermatophyta, Flowering Plants 8 (2): 31–76.

[S06] Stuckert, T. 1906. Segunda contribución al conocimiento de las gramináceas Argentinas. Anales del Museo Nacional de Buenos Aires, serie 3, 6: 409–555.

[WM09] Wang, H., M. J. Moore, P. S. Soltis, C. D. Bell, S. F. Brockington, R. Alexandre, C. C. Davis, M. Latvis, S. R. Manchester & D. E. Soltis. 2009. Rosid radiation and the rapid rise of angiosperm-dominated forests. Proceedings of the National Academy of Sciences of the USA 106 (10): 3853–3858.

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