Chacoan horned frog Ceratophrys cranwelli, copyright Alfredo Sabaliauskas.

Belongs within: Hyloidea.

The Ceratophryidae, South American horned frogs, are large-headed frogs found in arid habitats that are carnivorous as both adults and larvae.

Synapomorphies (from Frost et al. 2006): Transverse processes of anterior presacral vertebrae widely expanded; cranial bones dermosed; teeth fang-like, nonpedicellate; pars palatina of maxilla and premaxilla absent.

<==Ceratophryidae [Ceratophryinae, Ceratophryini]
| i. s.: Wawelia Casamiquela 1963 non Namysl. 1908 (ICBN)B00
| `--*W. gerholdi Casamiquela 1963B00
|--+--Lepidobatrachus Budgett 1899EJK08, FG06
| | |--L. asperEJK08
| | |--L. laevisFG06
| | `--L. llanensisFG06
| `--Chacophrys Reig & Limeses 1963EJK08, FG06
| `--C. pierottiiEJK08
`--Ceratophrys Wied-Neuwied 1824EJK08, FG06
| i. s.: C. joazeirensisFG06
| C. laevisP30
| C. testudoFG06
|--C. (Ceratophrys)C07
| |--C. (C.) auritaC07
| |--C. (C.) cranwelliC07
| `--C. (C.) ornataC07 [incl. C. prisca Ameghino 1899B00]
`--C. (Stombus Gravenhorst 1825)C07, FG06 [Stombinae]
|--C. (S.) stolzmanniC07
`--+--C. (S.) calcarataC07
`--C. (S.) cornutaC07

*Type species of generic name indicated


[B00] Báez, A. M. 2000. Tertiary anurans from South America. In: Heatwole, H., & R. L. Carroll (eds) Amphibian Biology vol. 4. Palaeontology. The evolutionary history of amphibians pp. 1388–1401. Surrey Beatty & Sons.

[C07] Cadle, J. E. 2007. The snake genus Sibynomorphus (Colubridae: Dipsadinae: Dipsadini) in Peru and Ecuador, with comments on the systematics of Dipsadini. Bulletin of the Museum of Comparative Zoology 158 (5): 183–284.

[EJK08] Evans, S. E., M. E. H. Jones & D. W. Krause. 2008. A giant frog with South American affinities from the Late Cretaceous of Madagascar. Proceedings of the National Academy of Sciences of the USA 105 (8): 2951–2956.

[FG06] Frost, D. R., T. Grant, J. Faivovich, R. H. Bain, A. Haas, C. F. B. Haddad, R. O. de Sá, A. Channing, M. Wilkinson, S. C. Donnellan, C. J. Raxworthy, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. M. Green & W. C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1–370.

[P30] Parker, H. W. 1930. Amphibia and Reptilia of the expedition to Brazil and Paraguay in 1926–27 by Dr. G. S. Carter and Mr. L. C. Beadle. Proceedings of the Linnean Society of London 141: 20.

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