Chlaenius

Chlaenius festivus, copyright Siga.

Belongs within: Oodini.

Chlaenius is a cosmopolitan genus of often metallic ground beetles with pubescent elytra that have the epipleura interrupted by a twisted fold near the apex. Members of the genus have been divided between over fifty subgenera and monophyly of the genus perhaps warrants investigation.

<==Chlaenius Bonelli 1810 [Chlaeniina]B12
    |--C. (Chlaenius) (see below for synonymy)BB01
    |    |--*C. (C.) festivus (Fabricius 1810) [=Carabus festivus]LM87
    |    |--C. (C.) aestivus Say 1823BB01
    |    |--C. (C.) augustus Newman 1838BB01
    |    |--C. (C.) cordicollis Kirby 1837BB01
    |    |--C. (C.) cursorBB01
    |    |--C. (C.) laticollisBB01
    |    |--C. (C.) platyderus Chaudoir 1856BB01
    |    |--C. (C.) ruficaudaBB01
    |    |--C. (C.) sericeus (Forster 1771)BB01
    |    |--C. (C.) solitariusBB01
    |    |--C. (C.) spoliatus [=Harpalus (C.) spoliatus]G20
    |    `--C. (C.) viduus Horn 1871BB01
    |--C. (Agostenus Motschulsky 1850) (see below for synonymy)BB01
    |    |--‘Agostenus’ glaucusBB01
    |    |--C. (A.) impunctifrons Say 1823BB01
    |    |--‘Agostenus’ lithophilusBB01
    |    |--‘Agostenus’ nemoralisBB01
    |    |--‘Agostenus’ pennsylvanicusBB01
    |    |--C. (A.) purpuricollis Randall 1838BB01
    |    |--C. (A.) tricolor Dejean 1826BB01
    |    |--‘Agostenus’ vaferBB01
    |    `--‘Agostenus’ variabilipesBB01
    |--C. (Anomoglossus Chaudoir 1856)BB01
    |--C. (Diapheropsophus)B66
    |    |--C. (D.) mellyiB66
    |    `--C. (D.) swinhoei Bates 1866B66
    |--C. (Epomis)B14
    |    |--C. (E.) circumscriptus (Duftschmid 1812)B14
    |    `--C. (E.) dejeaniiB14
    |--C. (Eurydactylus LaFerté Sénectère 1851) [incl. Glyptoderus LaFerté Sénectère 1851]BB01
    |    |--C. (E.) pimalicus Casey 1914BB01
    |    `--C. (E.) tomentosus (Say 1823)BB01
    |--C. (Lissauchenius) rufifemoralisOM08
    |    |--C. r. rufifemoralisOM08
    |    `--C. r. bimaculatus Dejean 1826OM08, LM87
    |--C. (Pelasmomimus Grundmann 1955)LM87
    |    `--C. (*P.) greyanus White 1841 [=C. grayianus]LM87
    `--C. (Pseudanomoglossus Bell 1960)BB01
         `--C. (P.) maxillosus Horn 1876BB01
Chlaenius incertae sedis:
  C. agrorumR26
  C. australis Dejean 1831LM87
  C. bengalensis Chaud. 1856B89
  C. cambodiensis Bates 1889B89
  C. chrysocephalusR26
  C. cinctusB89
  C. costiger Chaud. 1856B89
  C. darlingensis Castelnau 1867LM87
  C. flaviguttatus Macleay 1825 (see below for synonymy)LM87
  C. hamifer Chaudoir 1856 [incl. C. bihamatus Chaudoir 1856, C. queenslandicus Sloane 1910]LM87
  C. javanus Chaud. 1856 [incl. C. xanthopleurus]B89
  C. leucops [incl. C. aeruginosus Chaud. 1856]B89
  C. maculiger Castelnau 1867LM87
  C. nigricornis [incl. C. nigricornis var. melanocornis]C01
  C. nigripes Macleay 1886M86
  C. ophonoides Fairmaire 1843 (see below for synonymy)LM87
  C. planicornisB66
  C. rudicollis Chaudoir 1876 [incl. C. bimaculatus Macleay 1864 non Dejean 1826]LM87
  C. sinensisB89
  C. subcostatus Macleay 1864LM87
  C. tenellusG89
  C. velutinusR26
  C. virguliferC01
  C. xanthospilusB89

Chlaenius flaviguttatus Macleay 1825 [incl. Lissauchenius biguttatus Montrouzier 1860, C. binotatus Dejean 1826, C. guttatus Eschscholtz 1833, C. flaviguttatus immaculipennis Jedlicka 1951, C. maculifer Castelnau 1867, C. punctatus Chaudoir 1856, C. puncticeps Gemminger & Harold 1868]LM87

Chlaenius ophonoides Fairmaire 1843 [incl. C. peregrinus Chaudoir 1856, C. viridescens Gemminger & Harold 1868, C. viridis Montrouzier 1860 non Ménétries 1832]LM87

Chlaenius (Agostenus Motschulsky 1850) [inc. Brachylobus Chaudoir 1876, Chlaeniellus Reitter 1908, Pelasmus Motschulsky 1850]BB01

Chlaenius (Chlaenius) [incl. Callistometus Grundmann 1956, Hemichlaenius Lutschnik 1933 (preoc.), Lithochlaenius Kryzhanovskij 1976]BB01

*Type species of generic name indicated

References

[B12] Baehr, M. 2012. A new species of the genus Pheropsophus (Coleoptera: Carabidae: Brachininae) from northern Australia. Records of the Western Australian Museum 27 (1): 62–67.

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B66] Bates, H. W. 1866. On a collection of Coleoptera from Formosa sent home by R. Swinhoe, Esq., H. B. M. Consul, Formosa. Proceedings of the Zoological Society of London 1866: 339–355.

[B89] Bates, H.-W. 1889. Contributions a la faune Indo-Chinoise. 3e mémoire. Carabidae. Annales de la Société Entomologique de France, 6e série 9: 261–286.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, “Coleoptera”. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

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