Chordeumatida

Specimen belonging to the Australeuma mauriesi group, from Tasmanian Multipedes.

Belongs within: Eugnatha.
Contains: Neoatractosomatoidea, Anthroleucosomatidae, Cleidogonoidea, Diplomaragnidae, Conotylidae, Megalotylidae, Metopidiotrichinae.

The Chordeumatida are a group of relatively short, bristly millipedes.

Characters (from Hoffman 1990): Body with 26–30 segments in adults; ocelli usually present; gnathochilarium with short basal mentum and elongate lingual lamellae; ozopores absent; metaterga with well-defined median longitudinal suture; sterna not firmly attached to adjoining sternites but more or less flexibly articulated by connective tissue; males with coxal sacs in ninth and tenth pairs of legs only; gonopods composed of both pairs of legs of seventh segment.

<==Chordeumatida [Chordeumida]
    |  i. s.: NeoatractosomatoideaM03
    |         AnthroleucosomatidaeM03
    |         Nanogona Cook 1895H00
    |         Urochordeuma [Urochordeumatidae]H90
    |--ChordeumatideaS03b
    |    |  i. s.: ChordeumatidaeM03
    |    |           |--Chordeumella broelemanni (Verhoeff 1897)M03
    |    |           `--Chordeuma silvestre [incl. Craspedosoma nemasomoides, Cr. polydesmoides]B82
    |    `--CleidogonoideaH90
    |--StriariideaS03b
    |    |--Caseyidae [Caseyoidea]S03b
    |    |    |--CaseyaH90
    |    |    |--OpionaH90
    |    |    |--PlacernaH90
    |    |    |--VasingtonaH90
    |    |    `--ZantonaH90
    |    `--StriarioideaS03b
    |         |--Apterourus Loomis 1966 [Apterouridae]S03b
    |         |    |--*A. horizontalis Loomis 1966S03b
    |         |    `--A. serratus Shear 2003S03b
    |         `--+--RhiscosomidesH90 [RhiscosomididaeS03b]
    |            `--StriariidaeS03b
    |                 |--VaferiaH90
    |                 `--SpeostriariaH90
    |--+--DiplomaragnidaeS99
    |  `--Heterochordeumatoidea [Heterochordeumatidea]S03b
    |       |--HeterochordeumatidaeS02a
    |       |--Adrityla [Adritylidae]H90
    |       |--ConotylidaeH90
    |       |--MegalotylidaeS02b
    |       `--Metopidiotrichidae [Metopidiothricidae]S02a
    |            |--Schedotrigona Silvestri 1903 [Schedotrigonidae, Schedotrigoninae]S02a
    |            |--MetopidiotrichinaeS02a
    |            `--Neocambrisomatinae [Neocambrisomatidae]S02a
    |                 |--Neocambrisoma Mauriès 1987S02a
    |                 |--Nesiothrix Shear & Mesibov 1997S02a
    |                 |--Nipponothrix Shear & Tanabe 1994S02a
    |                 `--Australeuma Golovatch 1987S02a
    |                      `--A. mauriesi Shear & Mesibov 1997S02a
    |--CraspedosomatideaMG00
    |    |  i. s.: Craspedosoma [Craspedosomatidae, Craspedosomidae]MG00
    |    |           |--C. affine Koch & Berendt 1854RB93
    |    |           |--C. angulatum Koch & Berendt 1854RB93
    |    |           |--C. armatum (Gerstfeldt 1859) (n. d.) [=Julus armatus]MG00
    |    |           |--C. cylindricum Stuxberg 1876 (n. d.)MG00
    |    |           `--C. simileA99
    |    `--MastigophorophylloideaM03
    |         |--MastigophorophyllidaeM03
    |         |--HoffmaneumatidaeM03
    |         `--AltajellidaeMG00
    |              |--Elongeuma Golovatch 1982MG00
    |              |--Altajella Gulička 1972MG00
    |              |    `--*A. pallida Gulička 1972MG00
    |              `--Teleckophoron Gulička 1972MG00
    |                   `--*T. montanum Gulička 1972MG00
    `--Brannerioidea [Craspedosmatidea]SL03
         |--Niponiosoma [Niponiosomatidae]SH98
         |--TrachygonidaeSL03
         |--HeterolatzeliidaeSL03
         |--ChaemosomatidaeSL03
         |--Microlympia Shear & Leonard 2003 [Microlympiidae]SL03
         |    `--*M. echina Shear & Leonard 2003SL03
         |--Branneria Bollman 1893 [Branneriidae]S03a
         |    |--*B. carinata (Bollman 1888) [=Craspedosoma carinata]S03a
         |    `--B. bonoculus Shear 2003S03a
         |--TingupidaeSL03
         |    |--Tingupa Chamberlin 1910SL03
         |    |    `--T. pallidaSH98
         |    |--Buotus Chamberlin 1940SL03
         |    |    `--B. carolinusSH98
         |    `--Blancosoma Shear & Hubbard 1998SL03
         |         `--*B. scaturgo Shear & Hubbard 1998SH98
         `--KashmireumatidaeS02b
              |--KashmireumaS02b
              |    |--K. nepalensisS02b
              |    |--K. nielseniS02b
              |    `--K. schawallariS02b
              `--Vieteuma Golovatch 1984S02b
                   |--*V. topaliS02b
                   |--V. hubeiensis Mauriés & Nguyen-Duy Jaquemin 1997S02b
                   `--V. longi Shear 2002S02b

*Type species of generic name indicated

References

[A99] Ax, P. 1999. Das System der Metazoa II. Ein Lehrbuch der phylogenetischen Systematik. Gustav Fisher Verlag: Stuttgart (translated: 2000. Multicellular Animals: The phylogenetic system of the Metazoa vol. 2. Springer).

[B82] Berlese, A. 1882. Acari, Myriopoda et Scorpiones hucusque in Italia reperta. Fascicle 1. Typis Fratelli Salmin: Patavii.

[H00] Hannibal, J. T. 2000. Hexecontasoma, a new helminthomorph millipede (Hexecontasomatidae n. fam.) from the Mazon Creek, Illinois, fauna (Carboniferous, North America). Fragmenta Faunistica 43 (Suppl.): 19–35.

[H90] Hoffman, R. L. 1990. Diplopoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 835–860. John Wiley & Sones: New York.

[M03] Mauriès, J.-P. 2003. Schizmohetera olympica sp. n. from Greece, with a reclassification of the superfamily Neoatractosomatoidea (Diplopoda: Chordeumatida). Arthropoda Selecta 12: 9–16.

[MG00] Mikhaljova, E. V., & S. I. Golovatch. 2000. A review of the millipede fauna of Siberia (Diplopoda). Arthropoda Selecta 9: 103–118.

[RB93] Ross, A. J., & D. E. G. Briggs. 1993. Arthropoda (Euthycarcinoidea and Myriapoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 357–361. Chapman & Hall: London.

[S99] Shear, W. A. 1999. The milliped genus Diplomaragna confirmed for Taiwan, with the description of a new species (Diplopoda, Chordeumatida, Diplomaragnidae). Myriapodologica 6 (2): 11–18.

[S02a] Shear, W. A. 2002a. The millipede genus Metopidiothrix Attems (Diplopoda: Chordeumatida: Metopidiotrichidae). Invertebrate Systematics 16: 849–892.

[S02b] Shear, W. A. 2002b. Five new chordeumatidan millipeds from China: new species of Vieteuma (Kashmireumatidae) and Nepalella (Megalotylidae). Proceedings of the California Academy of Sciences 53 (6): 63–72.

[S03a] Shear, W. A. 2003a. Branneria bonoculus, n. sp., a second species in the North American milliped family Branneriidae (Diplopoda: Chordeumatida: Brannerioidea). Zootaxa 233: 1–7.

[S03b] Shear, W. A. 2003b. A second species of the rare milliped family Apterouridae (Diplopoda: Chordeumatida: Striarioidea). Proceedings of the Biological Society of Washington 116 (2): 509–514.

[SH98] Shear, W. A., & D. A. Hubbard, Jr. 1998. Cave millipeds of the United States. IV. A new genus and species from a high altitude cave in Colorado (Diplopoda, Chordeumatida, Tingupidae). Myriapodologica 5 (8): 85–94.

[SL03] Shear, W. A., & W. P. Leonard. 2003. Microlympiidae, a new milliped family from North America, and Microlympia echina, new genus and species (Diplopoda: Chordeumatida: Brannerioidea). Zootaxa 243: 1–11.

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