Clathurellidae

Clathurella rava, copyright Bestshells.

Belongs within: Conoidea.
Contains: Awateria, Scobinella, Glyphostoma (Glyphostoma), Lienardia (Lienardia), AcristaParaclathurella, Pleurotomoides, Nannodiella, Etrema (Etrema), Etrema (Etremopsis).

The Clathurellidae are a group of conoid gastropods with more or less fusiform shells, usually bearing strong, often cancellate sculpture. Most species possess a protoconch with a keel on at least part of the final whorl (Bouchet et al. 2011).

Characters (from Bouchet et al. 2011): Shell small- to medium-sized (5–40, usually 10–20 mm), fusiform to broadly fusiform, subsutural ramp usually unclearly separated. Sculpture mostly strong, usually cancellate, subsutural fold lacking. Shell surface often microgranular. Apertural armature often well developed, in form of pleats and denticles on both inner and outer lips; no true columellar pleats. Siphonal canal well expressed, moderately long. Anal sinus on subsutural ramp, deep, often (sub)tubular. Protoconch typically multispiral, up to six whorls, generally smooth with keeled last whorls; when paucispiral, protoconch usually smooth or spirally striated, sometimes with remaining carination in last portion. Operculum always absent. Radula of hypodermic marginal teeth, usually with relatively small solid basal part, sometimes with weak barb at tip. Distinct ligaments absent.

<==Clathurellidae [Clathurellinae, Defrancianina, Defranciinae]
    |--+--NannodiellaPK11
    |  `--Etrema Hedley 1918PK11, BD12
    |       |  i. s.: E. cratis Kilburn & Dekker 2008KD08
    |       |         E. obtusicostataH22
    |       |         E. opsimathes Melvill & Standen 1903H22
    |       |         ‘Pleurotrema’ polynesiensis Reeve 1845 [=Defrancia polynesiensis]KD08
    |       |         E. pyramidulaH22
    |       |         E. tenera (Hedley 1899) [=Glyphostoma aliciae var. tenera]KD08
    |       |--E. (Etrema)BK11
    |       |--E. (Etremopa Oyama 1953)BK11
    |       |    |--E. (*E.) subauriformis (Smith 1879) [=Drillia subauriformis]P66
    |       |    |--E. (E.) gainesii (Pilsbry 1895) [=Clathurella gainesii; incl. E. sawanensis Yokoyama 1927]P66
    |       |    |--E. (E.) nassoides (Reeve 1843)P66 (see below for synonymy)
    |       |    |--E. (E.) polydesma Hedley 1922P66
    |       |    |--E. (E.) saigoensis Makiyama 1927P66
    |       |    |--E. (E.) scalarina (Deshayes 1863) [=Pleurotoma scalarina]P66
    |       |    |--E. (E.) streptonotus (Pilsbry 1904) [=Drillia streptonotus]P66
    |       |    `--E. (E.) texta (Dunker 1860) [=Defrancia texta]P66
    |       `--E. (Etremopsis)BK11
    `--Clathurella Carpenter 1857PK11, BD12
         |--*C. rava (Hinds 1843) [=Clavatula rava]P66
         |--C. affinis Dall 1871P66
         |--C. arctata (Reeve 1846) [=Pleurotoma arctata]H09
         |--C. aurea Carpenter 1857P66
         |--C. balteata Pease 1860 [=Defrancia balteata]K65
         |--C. brunnea Pease 1860 [=Defrancia brunnea]K65
         |--C. buccinoides Pease 1860 [=Defrancia buccinoides]K65
         |--C. curtisiana (Hedley 1922)BK11
         |--C. cylindrica Pease 1860 [=Defrancia cylindrica]K65
         |--C. debilis (Hinds 1843) [=Clavatula debilis]H09
         |--C. donata (Hinds 1844) [=Clathurella donata]H22
         |--C. fuscomaculata Pease 1860 [=Defrancia fuscomaculata]K65
         |--C. hamiltoniH86
         |--C. harpa Pease 1860 [=Defrancia harpa]K65
         |--C. intercalaris (Carpenter 1856) [=Defrancia intercalaris]P66
         |--C. iodolabiata (Hornung & Mermod 1929) [=Mangilia (Clathurella) iodolabiata]KD08
         |--C. marshalliBW80
         |--C. nigrotincta (Montrouzier 1872)PK11
         |--C. paucicostata Pease 1860 [=Defrancia paucicostata]K65
         |--C. producta Pease 1860K65
         |--C. pulchella Pease 1860 [=Defrancia pulchella]K65
         |--C. purpureaH08 [=Mangilia purpureaH22; incl. C. reticosa Adams & Angas 1863H08]
         |--C. rigida (Hinds 1843) [=Clavatula rigida]P66
         |    |--C. r. rigidaP66
         |    `--C. r. fuscoligata (Carpenter 1856) [=Mangelia fuscoligata]P66
         |--C. rufinodis von Martens 1880H22
         |--C. serrata (Carpenter 1856) [=Defrancia serrata]P66
         `--C. thespesia (Melvill & Standen 1896) [=Daphnella thespesia]H09
Clathurellidae incertae sedis:
  Pleuropyramis Vredenburg 1921P66
    `--*P. quinqueangularis (Vredenburg 1921) [=Mangillia (Clathurella) quinqueangularis]P66
  Domenginella Vokes 1939P66
    `--*D. claytonensis (Gabb 1864) [=Turris claytonensis]P66
  Eoscobinella Powell 1942P66
    `--*E. tahuia Powell 1942P66
  Mitratoma Olsson 1930P66
    `--*M. bartschi (Olsson 1930) [=Scobinella (*Mitratoma) bartschi]P66
  Endiatoma Cossmann 1896P66
    `--*E. quadricincta (Cossmann 1883) [=Aphanitoma quadricincta]P66
  Nicolia Gregorio 1880P66
    `--*N. solitaria (Gregorio 1880) [=Borsonia (*Nicolia) solitaria]P66
  Plentaria Harris 1937P66
    |--*P. plenta (Aldrich & Harris 1895) [=Pleurotoma (Borsonia) plenta]P66
    `--P. horrida Harris 1937 [=P. plenta horrida]P66
  Cryptoborsonia Powell 1944P66
    |--*C. pleurotomella Powell 1944P66
    `--C. rugobela Powell 1944P66
  Protosurcula Casey 1904P66
    |--*P. gabbii (Conrad 1865) [=Surcula gabbii]P66
    |--P. platysoma (Heilprin 1880) [=Pleurotoma platysoma]P66
    `--P. tenuirostris Casey 1904P66
  Eucheilodon Gabb 1860P66
    |--*E. reticulata Gabb 1860P66
    |--E. crenocarinata Heilprin 1880P66
    |--E. gabbianum Casey 1904P66
    `--E. reticulatoides (Harris 1895) [=Pleurotoma (Eucheilodon) reticulatoides]P66
  Moniliopsis Conrad 1865P66
    |--*M. elaborata (Conrad 1833) [=Pleurotoma elaborata]P66
    |--M. hammetttensis (Harris 1937) [=Scobinella (Moniliopsis) hammettensis]P66
    |--M. nassiformis (Harris 1937) [=Scobinella (Moniliopsis) nassiformis]P66
    `--M. sculpturata (Aldrich 1911) [=Scobinella sculpturata]P66
  Glyptotoma Casey 1904P66
    |--*G. crassiplicata (Gabb 1860) [=Pleurotoma crassiplicata, Scobinella crassiplicata]P66
    |    |--G. c. crassiplicataP66
    |    `--G. c. montgomeryensis (Harris 1937) [=Bathytoma (Glyptotoma) montgomeryensis]P66
    |--G. conradiana (Aldrich 1895) [=Borsonia (Scobinella) conradiana]P66
    |--G. fisherana (Harris 1937) [=Bathytoma (Glyptotoma) fisherana]P66
    |--G. parvula Casey 1904P66
    `--G. rhombica Cooke 1928P66
  AwateriaP66
  ScobinellaP66
  Comarmondia Monterosato 1844BK11 (see below for synonymy)
    `--*C. gracilis (Montagu 1803) (see below for synonymy)P66
  Glyphostoma Gabb 1872BK11
    |  i. s.: G. cinerea [=Mangilia cinerea]H22
    |         G. conradiana Gabb 1869O27
    |         G. costicrenata (Cossmann 1900)P66
    |         G. cymodoce Dall 1919O27
    |         G. huberti Sowerby 1893H09
    |         G. polynesiense Reeve 1846H09
    |         G. rostrata Sysoev & Bouchet 2001BK11
    |         G. subcosticrenata MacNeil 1960P66
    |         G. trigonostoma Hervier 1897H22
    |--G. (Glyphostoma)BK11
    |--G. (Euglyphostoma Woodring 1970)BK11
    `--G. (Glyphostomops Bartsch 1934)BK11
         |--G. (*G.) hendersoni Bartsch 1934P66
         |--G. (G.) oenoa (Bartsch 1934) [=Glyphostomops oenoa]P66
         `--G. (G.) pinellasensis (Fargo 1953) [=Glyphostomops pinellasensis]P66
  Lienardia Jousseaume 1884BD12
    |  i. s.: L. bertiniana (Tapparone Canefri 1878) [=Clathurella bertiniana]H22
    |         L. brachyspiraH22
    |         L. chrysoleuca Melvill 1923 [incl. Glyphostoma permiscere Usticke 1969]BC01
    |         L. globulosa Hervier 1896BD12
    |         L. jousseaumei (Hervier 1896)BK11
    |         L. strombilla (Hervier 1895) [=Glyphostoma strombillum]H22
    |--L. (Lienardia)BK11
    `--L. (Thetidos Hedley 1899)BK11
         `--L. (*T.) morsura (Hedley 1899) [=*Thetidos morsura]P66
  Strombinoturris Hertlein & Strong 1951BK11
    `--*S. crockeri Hertlein & Strong 1951P66
  AcristaBK11
  Adanaclava Bartsch 1950BK11
    `--*A. adana Bartsch 1950 [=Crassispira (*Adanaclava) adana]P66
  Clathromangelia Monterosato 1884BK11
    |--C. quadrillum (Dujardin 1837) (see below for synonymy)P66
    |--C. clathrata (Marcel de Serres 1829) [=Pleurotoma clathrata]P66
    |--C. crebricostata (Carpenter 1865) [=Mangilia (Clathromangilia) crebricostata]O27
    |--C. duperrayi (Peyrot 1903) [=Mangilia duperrayi]P66
    |--C. interlirata (Stearns 1871) [=Mangilia (Clathromangilia) interlirata]O27
    |--C. levidensis (Carpenter 1865) [=Mangilia (Clathromangilia) levidensis]O27
    |--C. lineolata (Reeve 1846) [=Mangilia (Clathromangilia) lineolata]O27
    `--C. rhyssa (Dall 1919) [=Mangilia (Clathromangilia) rhyssa]O27
  Crockerella Hertlein & Strong 1951BK11
    |--*C. crystallina (Gabb 1865)P66 (see below for synonymy)
    |--C. hilli Hertlein & Strong 1951P66
    |--C. pederseni Hertlein & Strong 1951P66
    `--C. subdiaphana (Carpenter 1864)P66 [=Mangelia subdiaphanaP66, Cytharella subdiaphanaD19]
  Euclathurella Woodring 1928BK11
    |--*E. vendryesiana (Dall 1896) [=Clathurella vendryesiana]P66
    `--E. gertrudis (Toula 1909) [=Pleurotoma (Genota) gertrudis]P66
  ParaclathurellaBK11
  Pseudoetrema Oyama 1953BK11
    |--*P. fortilirata (Smith 1879)P66 [=Drillia fortilirataP66, Pseudorhaphitoma fortilirataH22]
    |--P. gerthi (Oostingh 1938) [=Clathurella gerthi]P66
    |--P. hukusimensis (Nomura & Zinbo 1935) [=Mangilia (Etrema) hukusimensis]P66
    |--P. ijzermani (Oostingh 1938) [=Clathurella ijzermani]P66
    |--P. kaloedanensis (Oostingh 1938) [=Clathurella kaloedanensis]P66
    `--P. sintikuensis (Nomura 1935) [=Lienardia (Etrema) sintikuensis]P66
  Turrella Laseron 1954BK11
    |--*T. tenuilirata (Angas 1871)P66 (see below for synonymy)
    |--T. asperrima Laseron 1954P66
    |--T. crassa Laseron 1954P66
    |--T. gracilis Laseron 1954P66
    |--T. granulosissima (Tenison-Woods 1879)P66 (see below for synonymy)
    |--T. letourneauxiana (Crosse & Fischer 1865) [=Pleurotoma letourneauxiana]P66
    |--T. morologus (Hedley 1922) [=Guraleus morologus]P66
    `--T. subcostata Laseron 1954P66
  PleurotomoidesP66

Clathromangelia quadrillum (Dujardin 1837) [=Pleurotoma quadrillum; incl. P. granum Philippi 1844, *Clathromangelia granum, P. rude Philippi 1836 non P. rudis Sowerby 1834]P66

Comarmondia Monterosato 1844BK11 [=Bellardia Bucquoy, Dautzenberg & Dollfus 1883 nec Allioni 1785 (ICBN) nec Robineau-Desvoidy 1863P66, Bellardiella Fischer 1883P66, Bellatula Strand 1929BK11]

*Comarmondia gracilis (Montagu 1803) [=Murex gracilis, *Bellardia gracilis, *Bellardiella gracilis, *Bellatula gracilis; incl. Fusus branscombi Clark 1849, Murex emarginatus Donovan 1804, M. oblongus Brocchi 1814]P66

*Crockerella crystallina (Gabb 1865)P66 [=Clathurella crystallinaP66, Philbertia crystallinaD19; incl. C. lowei Dall 1903P66]

Etrema (Etremopa) nassoides (Reeve 1843)P66 [=Pleurotoma nassoidesP66, Mangelia nassoidesH22; incl. Clathurella zonulata Angas 1867H22]

Turrella granulosissima (Tenison-Woods 1879)P66 [=Clathurella granulosissimaP66, Guraleus granulosissimusL54, Mangelia granulosissimaL54]

*Turrella tenuilirata (Angas 1871)P66 [=Clathurella tenuilirataP66, Defrancia tenuilirataH22, Guraleus tenuiliratusL54, Mangelia tenuilirataH22, Pseudoraphitoma tenuilirataL54, Turrella multilirata (l. c.)L54]

*Type species of generic name indicated

References

[BD12] Bandel, K., & D. T. Dockery III. 2012. Protoconch characters of Late Cretaceous Latrogastropoda (Neogastropoda and Neomesogastropoda) as an aid in the reconstruction of the phylogeny of the Neogastropoda. Freiberger Forschungshefte C542 (psf 20): 93–128.

[BK11] Bouchet, P., Y. I. Kantor, A. Sysoev & N. Puillandre. 2011. A new operational classification of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 273–308.

[BW80] Bouchet, P., & A. Waren. 1980. Revision of the north-east Atlantic bathyal and abyssal Turridae (Mollusca, Gastropoda). Journal of Molluscan Studies Supplement 8: 1–119.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[D19] Dall, W. H. 1919. Descriptions of new species of mollusks of the family Turritidae from the west coast of America and adjacent regions. Proceedings of the United States National Museum 56 (2288): 1–86, pls 1–24.

[H08] Hedley, C. 1908. Studies on Australian Mollusca. Part X. Proceedings of the Linnean Society of New South Wales 33: 456–489, pls 7–10.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H22] Hedley, C. 1922. A revision of the Australian Turridae. Records of the Australian Museum 13 (6): 213–359, pls 42–56.

[H86] Hutton, F. W. 1886. The Mollusca of the Pareora and Oamaru systems of New Zealand. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 205–237.

[K65] Kay, E. A. 1965. Marine molluscs in the Cuming collection, British Museum (Natural History) described by William Harper Pease. Bulletin of the British Museum (Natural History): Zoology Supplement 1: 1–96, 14 pls.

[KD08] Kilburn, R. N., & H. Dekker. 2008. New species of turrid conoideans (Gastropoda: Conoidea) from the Red Sea and Arabia. Basteria 72 (1–3): 1–19.

[L54] Laseron, C. F. 1954. Revision of the New South Wales Turridae (Mollusca). Royal Zoological Society of New South Wales: Sydney.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 1. Stanford University Press: Stanford University (California).

[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: an evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.

[PK11] Puillandre, N., Y. I. Kantor, A. Sysoev, A. Couloux, C. Meyer, T. Rawlings, J. A. Todd & P. Bouchet. 2011. The dragon tamed? A molecular phylogeny of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 259–272.

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