Cleoniceras besairei, from Antonov.

Belongs within: Pachydiscidae.

The Cleoniceratidae are a Lower Cretaceous (Lower Albian) lineage of ammonites (Wright et al. 1996).

Characters (from Wright et al. 1996): Round-whorled to compressed and high-whorled; venter rounded, subtabulate, or acute; ribs Y-shaped to falcoid; primaries single or branching at umbilical edge, with or without umbilical bullae, branching again at midflank or beyond; some forms secondarily smooth; constrictions present throughout or absent, commonly several close together before aperture. Microconchs small, commonly with venter of body chamber broadening and coiling inward; macroconchs rather large, with ornament weakening on outer whorl. Suture with wide, more or less asymmetrical L and up to six auxiliary lobes in high-whorled forms.

<==Cleoniceratidae [Cleoniceratinae, Lemuroceratinae, Vnigriceratinae]WCH96
    |  i. s.: Pachygrycia Jeletzky & Stelck 1981WCH96
    |           `--*P. canadensis Jeletzky & Stelck 1981WCH96
    |         Colvillia Imlay 1961WCH96
    |           |--*C. kenti Imlay 1961WCH96
    |           `--C. crassicostataWCH96
    |--Freboldiceras Imlay 1959WCH96
    |    `--*F. singulare Imlay 1959WCH96
    |--+--Cymahoplites Spath 1922 [=Vjasemskiceras Sazonova 1961]WCH96
    |  |    `--*C. kerenskianus (Bogoslovskii 1902) [=Ammonites kerenskianus]WCH96
    |  |--Lemuroceras Spath 1942WCH96
    |  |    |--*L. aburense (Spath 1933) [=Pseudohaploceras aburense]WCH96
    |  |    `--L. spathiP93
    |  `--Moretella Collignon 1963WCH96
    |       `--*M. madagascariensis (Collignon 1949) [=Pseudosonneratia madagascariensis]WCH96
    |--+--Arcthoplites Spath 1925 [incl. Bellidiscus Saveliev 1973, Subarcthoplites Casey 1954]WCH96
    |  |    |--*A. jachromensis (Nikitin 1888) [=Ammonites jachromensis]WCH96
    |  |    |--A. belli (McLearn 1945) [=Lemuroceras belli, *Subarcthoplites belli]WCH96
    |  |    `--A. probus (Saveliev 1973) [=*Bellidiscus probus]WCH96
    |  `--+--Tetrahoplitoides Casey 1954 [=Coloboceras Crickmay 1927 non Trouessart 1889]WCH96
    |     |    `--*T. stantoni (Anderson 1902) [=Sonneratia stantoni, *Coloboceras stantoni]WCH96
    |     |--Brewericeras Casey 1954WCH96
    |     |    |--*B. breweri (Gabb 1864) [=Ammonites breweri]WCH96
    |     |    `--B. hulenenseWCH96
    |     `--Leconteites Casey 1954 (see below for synonymy)WCH96
    |          |--*L. lecontei (Anderson 1902) [=Desmoceras lecontei]WCH96
    |          |--*Vnigriceras’ emendatum Saveliev 1973WCH96
    |          |--‘Vnigriceras (*Astrodiscus)’ insegestum Saveliev 1973WCH96
    |          `--L. sacramenticus (Anderson 1902) [=Pachydiscus sacramenticus, *Puzosigella sacramentica]WCH96
    `--+--Anadesmoceras Casey 1954 [incl. Carloscaceresiceras Etayo Serna 1979]WCH96
       |    |--*A. strangulatum Casey 1954WCH96
       |    |--*Carloscaceresiceras’ caceresi Etayo Serna 1979WCH96
       |    `--A. emendatumWCH96
       `--Cleoniceras Parona & Bonarelli 1897WCH96
            |--*C. (Cleoniceras) cleon (Orbigny 1850) [=Ammonites cleon]WCH96
            |--C. (Anacleoniceras Mirzoev 1969) [incl. Cleonella Destombes 1970 (n. n.)]WCH96
            |    |--C. (*A.) caseyi (Mirzoev 1969) [=*Anacleoniceras caseyi]WCH96
            |    `--C. dimorphum Casey 1966 [=*Cleonella dimorpha]WCH96
            |--C. (Eocleoniceras Saveliev 1992)WCH96
            |    `--C. (*E.) remotum Saveliev 1992WCH96
            |--C. (Grycia Imlay 1961) [incl. Paracleoniceras Collignon 1963]WCH96
            |    |--C. (*G.) sablei Imlay 1961WCH96
            |    `--C. (G.) besairei Collignon 1949 [=*Paracleoniceras besairei]WCH96
            `--C. (Neosaynella Casey 1954)WCH96
                 `--C. (*N.) inornatum Casey 1954WCH96

Leconteites Casey 1954 [incl. Astrodiscus Saveliev 1973 nec Ludwig 1866 nec Schulze 1875, Puzosigella Casey 1954, Vnigriceras Saveliev 1973]WCH96

*Type species of generic name indicated


[P93] Page, K. N. 1993. Mollusca: Cephalopoda (Ammonoidea: Phylloceratina, Lytoceratina, Ammonitina and Ancyloceratina). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 213–227. Chapman & Hall: London.

[WCH96] Wright, C. W., J. H. Calloman & M. K. Howarth. 1996. Treatise on Invertebrate Paleontology pt L. Mollusca 4, revised vol. 4. Cretaceous Ammonoidea. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

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