Cleroidea

Phycosecis litoralis, copyright Udo Schmidt.

Belongs within: Coleoptera.
Contains: Byturidae, Peltidae, TrogossitidaeCleridae, Melyridae.

The Cleroidea are a clade of beetles supported by molecular data, as well as by the structure of the aedeagus in males (McKenna et al. 2015). Representatives include the Prionoceridae, an Old World family of elongate beetles with soft elytra, pentamerous tarsi and, in males, a comb on the inner edge of the fifth fore-tarsal segment. Phycosecis is an Australasian genus of ovate beetles with the head partially concealed by a projection of the pronotum and with a single-segmented club on the antenna.

<==Cleroidea [Melyrides]MW15
    |--RentoniinaeMW15
    |    |--RentonellumMW15
    |    `--Parentonium australicumLB91, B74
    `--+--ByturidaeMW15
       `--+--Phloiophilus [Phloiophilidae, Phloiophilinae]RS15
          |    `--P. edwardsii Stephens 1830B14
          `--+--+--+--Metaxina [Metaxinidae]MW15
             |  |  `--ChaetosomatidaeRS15
             |  |       |--Chaetosoma scaritides Westwood 1851MW15, B14
             |  |       |--ChaetosomodesB14
             |  |       `--MalgassochaetusB14
             |  `--Ostoma Laicharting 1781RS15, L02 [incl. Gaurambe Thomson 1859L02; Ostomatidae, Ostomidae]
             |       |--O. ferrugineumRS15
             |       `--O. pudicum Oll. 1889M96
             `--+--+--PeltidaeRS15
                |  `--+--TrogossitidaeRS15
                |     `--ThanerocleridaeMW15
                |          |  i. s.: Ababa Casey 1897 (see below for synonymy)O02
                |          |           `--A. tantilla [incl. *A. crinita Casey 1897]O02
                |          |--Zenodosus Wolcott 1910 [Zenodosinae]B14
                |          |    `--*Z. sanguineus (Say 1835) [=Clerus sanguineus]O02
                |          `--ThaneroclerinaeMW15
                |               |--IsoclerusMW15
                |               `--Thaneroclerus Chapin 1924 (see below for synonymy)O02
                |                    |--*T. buquet (Lefebvré 1835)O02 (see below for synonymy)
                |                    |--T. ainoB14
                |                    |--T. impressusB14
                |                    `--T. termitincolaB14
                `--+--CleridaeRS15
                   `--+--Larinotus [Larinotinae]RS15
                      |    `--L. umbilicatusRS15
                      `--+--Phycosecis [Phycosecidae]MW15
                         |    |--P. algarum Pascoe 1875M86
                         |    |--P. ammophilusB14
                         |    |--P. atomariaB14
                         |    |--P. discoideaB14
                         |    |--P. hilliB14
                         |    |--P. limbata (Fabricius 1781)B14
                         |    `--P. litoralis Pascoe 1875M86
                         `--+--MelyridaeRS15
                            `--PrionoceridaeMW15
                                 |--PrionocerusRS15
                                 |    |--P. bicolor Redtenbacher 1868B14
                                 |    `--P. coeruleipennisB14
                                 `--IdgiaMW15
                                      |--I. flavicollisF89
                                      |--I. flavirostrisF89
                                      |--I. moupinensis Fairmaire 1889F89
                                      `--I. oculataF89
Cleroidea incertae sedis:
  MauroniscidaeMW15
    |--Amecomycter Majer 1995B14, M02
    |--Mauroniscus Bourgeois 1911M02
    |    |--M. boliviensisB14
    |    `--M. maculatus Pic 1927B14
    |--Mectemycor Majer 1995B14, M02
    |    |--M. linearis (Fall 1930)M02
    |    |--M. sericeus Majer 1995M02
    |    `--M. strangulatus Majer 1995M02
    |--Mecomycter Horn 1882B14, M02
    |    |--M. majeri Howell 1997M02
    |    |--M. omalinus Horn 1882M02
    |    `--M. testaceous Majer 1995M02
    `--Scuromanius Majer 1995B14, M02
         |--S. facetus (Casey 1895)M02
         |--S. liebecki (Blaisdell 1929)M02
         `--S. wickhami Majer 1995M02
  Rhadalidae [Haplocneminae, Rhadalinae]MW15
    |--AnthriboclerusM02
    |--Aplocnemus Stephens 1830M02
    |--Indiodasytes Pic 1916M02
    |--Aploceble Majer 1998M02
    |--Eucymbolus Champion 1913M02
    |    `--E. cyaneus Champion 1913M02
    |--Dasyrhadus Fall 1910M02
    |    |--D. impressicollis Fall 1910M02
    |    `--D. longior Fall 1910M02
    |--Rhadalus LeConte 1852 [incl. Cymbolus Gorham 1886]M02
    |    |--R. lecontei Casey 1895M02
    |    `--R. testaceous LeConte 1852M02
    `--Semijulistus Schilsky 1894 [incl. Celsus Lewis 1895, Eurelymis Casey 1895]M02
         |--S. bicoloripes Pic 1928M02
         `--S. rubrithorax Pic 1928M02

Ababa Casey 1897 [incl. Cleridopsis Champion 1913, Prionodera Wolcott 1910, Prionostichaeus Wolcott 1911, Wolcottella Lucas 1920]O02

Thaneroclerus Chapin 1924 [incl. Thanatereroclerus Gemminger & Harold 1869, Thaneclerus Desmarest 1860]O02

*Thaneroclerus buquet (Lefebvré 1835)O02 [=Clerus buquetO02; incl. T. girodi Chv. 1880FS90, T. sanguinolentusFS90]

*Type species of generic name indicated

References

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[B74] Britton, E. B. 1974. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 62–89. Melbourne University Press.

[F89] Fairmaire, L. 1889. Coléoptères de l’intérieur de la Chine. 5e partie. Annales de la Société Entomologique de France, 6e série 9: 5–84.

[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.

[L02] Leschen, R. A. B. 2002. Trogossitidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 263–266. CRC Press.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

[M96] Masters, G. 1896. Catalogue of the described Coleoptera of Australia. Supplement, part II. Dytiscidae, Gyrinidae, Hydrophyllidae, Staphylinidae, Pselaphidae, Paussidae, Silphidae, Scaphididae, Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydiidae, Cucujidae, Cryptophagidae, Lathridiidae, Mycetophagidae, Dermestidae, Byrrhidae, Parnidae, Heteroceridae. Proceedings of the Linnean Society of New South Wales 21 (Suppl.): 695–754.

[M02] Mayor, A. J. 2002. Melyridae Leach 1815. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 281–304. CRC Press.

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[O02] Opitz, W. 2002. Cleridae Latreille 1804. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 267–280. CRC Press.

[RS15] Robertson, J. A., A. Ślipiński, M. Moulton, F. W. Shockley, A. Giorgi, N. P. Lord, D. D. McKenna, W. Tomaszewska, J. Forrester, K. B. Miller, M. F. Whiting & J. V. McHugh. 2015. Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia). Systematic Entomology 40: 745–778.

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