Bank voles Clethrionomys glareolus, photographed by Plantsurfer.

Belongs within: Arvicolinae.

Clethrionomys, the red-backed voles, is a Holarctic genus of small rodents in which most species (as the vernacular name indicates) have reddish fur dorsally, with the remainder of the fur being grey.

<==Clethrionomys Tilesius 1850RR75
    |--C. centralisIT07
    |--C. fraterRR75
    |--C. gapperi (Vigors 1830)RR75
    |    |--C. g. gapperiRR75
    |    |--C. g. athabascaeRR75
    |    |--C. g. brevicaudusHC52
    |    |--C. g. galeiRR75
    |    |--C. g. loringi (Bailey 1897) [=Evotomys gapperi loringi]B75
    |    |--C. g. nivarius (Bailey 1897)RR75, HC52 [=Evotomys nivariusHC52, C. occidentalis nivariusRR75, E. gapperi nivariusHC52]
    |    |--C. g. pallescens Hall & Cockrum 1952 (see below for synonymy)HC52
    |    |--C. g. phaeus (Swarth 1911) [=Evotomys phaeus]HC52
    |    |--C. g. saturatusHC52
    |    |--C. g. solus Hall & Cockrum 1952HC52
    |    |--C. g. stikinensis Hall & Cockrum 1952HC52
    |    `--C. g. wrangeli (Bailey 1897) [=Evotomys wrangeli]HC52
    |--C. glareolus (Schreber 1780)FK55 [incl. Lemmus arvalisE38, Arvicola rufescens de Selys Longchamps 1836HC52]
    |--C. montanus Imaizumi 1972I92
    |--C. occidentalis (Merriam 1890) [=Evotomys occidentalis]HC52
    |    |--C. o. occidentalis (see below for synonymy)HC52
    |    |--C. o. californicus (Merriam 1890) [=Evotomys californicus]HC52
    |    |--C. o. caurinus (Bailey 1898) [=Evotomys caurinus, C. gapperi caurinus]HC52
    |    |--C. o. mazama (Merriam 1897) [=Evotomys mazama, C. californicus mazama]HC52
    |    `--C. o. obscurus (Merriam 1897) [=Evotomys obscurus, C. californicus obscurus, C. mazama obscurus]HC52
    |--C. rex Imaizumi 1971I92
    |--C. rufocanusRR75
    |    |--C. r. rufocanusRR75
    |    `--C. r. bedfordiaeRR75
    |--C. rutilus (Pallas 1779) [=Evotomys rutilus]RR75
    |    |--C. r. rutilusRR75
    |    |--C. r. albiventer Hall & Gilmore 1932RR75
    |    |--C. r. dawsoni (Merriam 1888) [=Evotomys dawsoni; incl. E. alascensis Miller 1898]RR75
    |    `--C. r. mikadoRR75
    |--C. sikotanensisIT07
    `--C. yesomontanus Kishida 1931TYM08

Clethrionomys gapperi pallescens Hall & Cockrum 1952 [=C. gapperi rufescens Smith 1940 non Arvicola rufescens de Selys Longchamps 1836]HC52

Clethrionomys occidentalis occidentalis (Merriam 1890) [=C. californicus occidentalis, Evotomys gapperi occidentalis; incl. E. pygmaeus Rhoads 1894]HC52

*Type species of generic name indicated


[B75] Bowles, J. B. 1975. Distribution and biogeography of mammals of Iowa. Special Publications, The Museum, Texas Tech University 9: 1–184.

[E38] Ewing, H. E. 1938. North American mites of the subfamily Myobiinae, new subfamily (Arachnida). Proceedings of the Entomological Society of Washington 40 (7): 180–197.

[FK55] Felten, H., & C. König. 1955. Einige Säugetiere aus dem Zentralmassiv, Südfrankreich. Senckenbergiana Biologica 36: 267–269.

[HC52] Hall, E. R., & E. L. Cockrum. 1952. Comments on the taxonomy and geographic distribution of North American microtines. University of Kansas Publications, Museum of Natural History 5 (23): 293–312.

[IT07] Isaac, N. J. B., S. T. Turvey, B. Collen, C. Waterman & J. E. M. Baillie. 2007. Mammals on the EDGE: conservation priorities based on threat and phylogeny. PloS One 2 (3): e296.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[RR75] Rausch, R. L., & V. R. Rausch. 1975. Relationships of the red-backed vole, Clethrionomys rutilus (Pallas), in North America: karyotypes of the subspecies dawsoni and albiventer. Systematic Zoology 24 (2): 163–170.

[TYM08] Tennent, W. J., M. Yasuda & K. Morimoto. 2008. Lansania Journal of arachnology and zoology—a rare and obscure Japanese natural history journal. Archives of Natural History 35 (2): 252–280.

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