Clivina

Clivina vagans, copyright Birgit E. Rhode.

Belongs within: Scaritinae.

Clivina is a cosmopolitan genus of ground beetles with a sharp spine externally on the mid-tibia above or at the apex, and a concavity on either side of the peduncle.

<==Clivina Latreille 1802 [incl. Ceratoglossa Macleay 1863, Scolyptus Putzeys 1863]LM87
    |--C. (Clivina) [incl. Reichardtula Whitehead in Reichardt 1977]BB01
    |    |--C. (C.) fossor (Linnaeus 1758)BB01 (see below for synonymy)
    |    |--C. (C.) collaris (Herbst 1784)BB01
    |    `--C. (C.) impressefrons LeConte 1844BB01
    |--C. americana Dejean 1831BB01
    |--C. (Paraclivina Kult 1947)BB01
    |    `--C. (P.) bipustulata (Fabricius 1801)BB01
    `--C. (Semiclivina Kult 1947)BB01
         |--C. (S.) dentipes Dejean 1825BB01
         `--C. (S.) vespertina Putzeys 1866BB01
Clivina incertae sedis:
  C. agonaB89b
  C. atrata Putzeys 1863 [incl. C. obliterata Sloane 1896]LM87
  C. atridorsis Sloane 1905LM87
  C. attenuata [incl. C. melanaria, C. picipes]B89b
  C. bacillaria Bates 1889B89b
  C. banksi Sloane 1907LM87
  C. biplagiata Putzeys 1866LM87
  C. blackburni Sloane 1896LM87
  C. bovillae Blackburn 1890LM87
  C. brevisterna Sloane 1916 [incl. C. brevisterna major Sloane 1917]LM87
  C. brunnicolor Sloane 1916LM87
  C. burgeoniB55
  C. cava Putzeys 1866LM87
  C. convexicollisB89b
  C. coronata Putzeys 1873LM87
  C. cribriforns Sloane 1905LM87
  C. curvidens LaFerté 1850 [incl. C. microphthalmus Putzeys 1863, *Scolyptus microphthalmus]LM87
  C. cylindripennis Sloane 1905LM87
  C. dampieri Sloane 1916LM87
  C. darwini Sloane 1916LM87
  C. diluta Darlington 1953LM87
  C. dingo Sloane 1905LM87
  C. diophthalmica Basilewsky 1955B55
  C. doddi Sloane 1905LM87
  C. dumoliniB55
  C. exilis Sloane 1916LM87
  C. froggatti Sloane 1896LM87
  C. gibbaR26
  C. grandiceps Sloane 1896LM87
  C. grata Darlington 1953LM87
  C. hackeri Sloane 1907LM87
  C. helferiB89b
  C. indica Putz. 1846B89b
  C. inopaca Darlington 1962LM87
  C. kershawi Sloane 1916LM87
  C. lataB89b
  C. machadoi Basilewsky 1955B55
  C. marginipennis Putz. 1846FS90
  C. ‘microphthalma’ Burgeon 1937 non C. microphthalmus Putzeys 1863B55
  C. misella Sloane 1905LM87
  C. mordaxB89b
  C. natalensisB55
  C. nigra Sloane 1905LM87
  C. obliquicollis Sloane 1905LM87
  C. oblongicollis Putzeys 1863BM87
  C. olliffi Sloane 1896LM87
  C. ovalipennis Sloane 1905 [=C. ovipennis Sloane 1896 non Chaudoir 1846]LM87
  C. pallidiceps Sloane 1905LM87
  C. parryi Putzeys 1863 [incl. Eupalamus clivinoides]B89a
  C. pectonoda Sloane 1905LM87
  C. physopleura Burgeon 1935B55
  C. planifrons Sloane 1907LM87
  C. robusta Sloane 1905LM87 (see below for synonymy)
  C. rugithoraxM83
  C. scripta Putz 1846E12
  C. sculpticeps Darlington 1953LM87
  C. siamicaB89b
  C. sloanei Csiki 1927 [=C. femoralis Sloane 1916 non Putzeys 1846]LM87
  C. sulcaticeps Sloane 1923LM87
  C. sulcicollis Sloane 1896 [=C. punctaticeps sulcicollis]LM87
  C. sulcigeraB89b
  C. thoracicaR26
  C. transversaB89b
  C. wiluna Darlington 1953LM87
  C. zebi Kult 1951LM87
  C. australasiae groupS96
    |--C. pectoralis Putzeys 1868S96, LM87
    |--C. australasiae sectionS96
    |    |--C. australasiae Boheman 1858LM87
    |    |--C. helmsi Blackburn 1892S96, LM87
    |    |--C. leai Sloane 1896S96, LM87 [incl. C. leai apicalis Sloane 1896LM87]
    |    |--C. queenslandica Sloane 1896S96, LM87
    |    `--C. vittata Sloane 1896S96, LM87
    |--C. basalis sectionS96
    |    |--C. angustipes Putzeys 1868S96, LM87
    |    |--C. basalis Chaudoir 1843S96, LM87 (see below for synonymy)
    |    |--C. dilutipes Putzeys 1868S96, LM87 (see below for synonymy)
    |    |--C. isogona Putzeys 1868S96, LM87
    |    |--C. lepida Putzeys 1866S96, LM87 [incl. C. juvensis Putzeys 1866LM87]
    |    |--C. rubripes Putzeys 1868S96, LM87
    |    |--C. simulans Sloane 1896S96, LM87
    |    `--C. vagans Putzeys 1866S96, LM87 [incl. C. lepida tasmaniensis Sloane 1896LM87]
    `--C. sellata sectionS96
         |--C. aequalis Blackburn 1890S96, LM87
         |--C. dimidiata Putzeys 1866S96, LM87 [incl. C. melanopyga Putzeys 1866LM87]
         |--C. ferruginea Putzeys 1868S96, LM87
         |--C. nana Sloane 1896S96, LM87
         |--C. occulta Sloane 1896S96, LM87
         |--C. sellata Putzeys 1866S96, LM87 [incl. C. sellata inconspicua Sloane 1896LM87]
         `--C. suturalis Putzeys 1863S96, LM87 [incl. C. dorsalis Blackburn 1890LM87, C. verticalis Putzeys 1866LM87]
  C. cribrosa groupS96
    |--C. boops Blackburn 1890S96, LM87
    |--C. cribrosa Putzeys 1868LM87
    |--C. fortis Sloane 1896S96, LM87
    `--C. frenchi Sloane 1896S96, LM87
  C. heterogena groupS96
    |--C. australica Sloane 1896S96, LM87
    |--C. difformis Putzeys 1868S96, LM87
    |--C. flava Putzeys 1868S96, LM87
    |--C. heterogena Putzeys 1866 [incl. C. angustula Putzeys 1866, C. deplanata Putzeys 1866]LM87
    |--C. odontomera Putzeys 1868S96, LM87
    |--C. oodnadattae Blackburn 1894S96, LM87
    `--C. tuberculifrons Blackburn 1890S96, LM87
  C. obliquata groupS96
    |--C. bitincta Sloane 1905LM87 [=C. bicolor Sloane 1896 non Putzeys 1846S96, LM87]
    |--C. cylindriformis Sloane 1896S96, LM87
    |--C. debilis Blackburn 1890S96, LM87
    |--C. denticollis Sloane 1896S96, LM87 [incl. C. eyrensis Blackburn 1900LM87]
    |--C. eremicola Blackburn 1894S96, LM87
    |--C. obliquata Putzeys 1866 [incl. C. adelaidae Blackburn 1890]LM87
    |--C. obsoleta Sloane 1896S96, LM87
    |--C. riverinae Sloane 1896S96, LM87
    `--C. wildi Blackburn 1890S96, LM87
  C. planiceps groupS96
    |--C. carpentaria Sloane 1896S96, LM87
    |--C. planiceps (Putzeys 1863)LM87 (see below for synonymy)
    `--C. quadratifrons Sloane 1896S96, LM87
  C. procera groupS96
    |--C. elegans Putzeys 1863S96, LM87
    |--C. gracilipes Sloane 1896S96, LM87
    |--C. macleayi Sloane 1896S96, LM87
    |--C. marginata (Putzeys 1868)S96, LM87 [=Scolyptus marginatusLM87]
    |--C. mastersi Sloane 1896S96, LM87
    |--C. monilicornis Sloane 1896S96, LM87
    |--C. nyctosyloides Putzeys 1868S96, LM87 [incl. C. interstitialis Sloane 1896LM87]
    |--C. oblonga (Putzeys 1873)S96, LM87 (see below for synonymy)
    |--C. obscuripes (Blackburn 1890)S96, LM87 [=Scolyptus obscuripesLM87]
    |--C. procera Putzeys 1866 [=Scolyptus procerus]S96
    |--C. prominens Putzeys 1866 [=Scolyptus prominens]S96
    `--C. regularis Sloane 1896S96, LM87
  C. punctaticeps groupS96
    |--C. emarginata Putzeys 1868S96
    |--C. lobipes Sloane 1896S96, LM87
    |--C. punctaticeps Putzeys 1868LM87
    `--C. tumidipes Sloane 1896S96, LM87

Clivina basalis Chaudoir 1843S96, LM87 [incl. C. ephippiata Putzeys 1846LM87, C. eximia Sloane 1896LM87, C. felix Sloane 1896LM87, C. microdon Putzeys 1866LM87]

Clivina dilutipes Putzeys 1868S96, LM87 [incl. C. sydneyensis Sloane 1896LM87, C. dilutipes victoriae Sloane 1905LM87]

Clivina oblonga (Putzeys 1873)S96, LM87 [=Scolyptus oblongusLM87; incl. S. abbreviatus Putzeys 1873LM87, Clivina abbreviataS96]

Clivina planiceps (Putzeys 1863)LM87 [=Scolyptus planicepsLM87; incl. S. crassicollis Putzeys 1866LM87, Clivina crassicollisS96, Ceratoglossa rugiceps Macleay 1863LM87, Scolyptus rugicepsB90]

Clivina robusta Sloane 1905LM87 [incl. Ceratoglossa foveiceps Macleay 1863 non Cl. foveiceps Putzeys 1845LM87, Scolyptus foveicepsB90]

Clivina (Clivina) fossor (Linnaeus 1758)BB01 [=Tenebrio fossorLM87; incl. Scarites arenarius Fabricius 1792LM87, *Clivina arenariaLM87]

*Type species of generic name indicated

References

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B89a] Bates, H. W. 1889a. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 100–111.

[B89b] Bates, H.-W. 1889b. Contributions a la faune Indo-Chinoise. 3e mémoire. Carabidae. Annales de la Société Entomologique de France, 6e série 9: 261–286.

[B90] Blackburn, T. 1890. Notes on Australian Coleoptera, with descriptions of new species. Part V. Proceedings of the Linnean Society of New South Wales, series 2, 4 (4): 1247–1276.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1–98.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

[S96] Sloane, T. G. 1896. On the Australian Clivinides (fam. Carabidae). (Revision of the Australian species of the genus Clivina with the description of a new genus, Clivinarchus). Proceedings of the Linnean Society of New South Wales 21 (2): 143–257.

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