Reed sac spider Clubiona phragmites, from Pavouci.

Belongs within: Clubionoidea.

Clubiona is a large, cosmopolitan genus of sac spiders in which the fourth pair of legs is longer than the remainder. Members of the genus are nocturnal hunters on vegetation, sheltering in rolled leaves or under stones or leaf litter during the day (Chen & Huang 2012).

Characters (from Chen & Huang 2012): Yellow or brownish; carapace concave and gradually narrowing towards cephalic region; thoracic groove distinct; abdomen cylindrical. Eyes homogeneous, arranged in two parallel transverse rows occupying most of head width; posterior eye row straight or slightly procurved, wider than anterior row. Male palp with retrolateral tibial apophysis; retrobasal cymbial spur absent. Legs prograde; tarsi with two claws; leg I as long as or shorter than leg II, leg IV longest.

Clubiona Latreille 1804CH12
|--*C. pallidula (Clerck 1757)CH12 [incl. Aranea holosericea Linnaeus 1758C01, Clubiona holosericeaC01]
|--C. alascensis Marx 1891S06
|--C. aloma [=Aranea aloma]C01
|--C. amarantha [=Aranea amarantha]C01
|--C. arctica Marx 1891S06
|--C. asrevida Ono 1992CH12
|--C. atra Risso 1826R26
|--C. bandoi Hayashi 1995CH12
|--C. barbara Lucas 1846E12
|--C. blesti Forster 1979PVD10
|--C. borealis Thorell 1870S06
|--C. brevipesV09
|--C. brevispina Huang & Chen in Chen & Huang 2012CH12
|--C. cada Forster 1979PVD10
|--C. cambridgei Koch 1873PVD10
|--C. canadensis Emerton 1890PS08
|--C. chathamensis Simon 1905PVD10
|--C. cirrosa Ono 1989CH12
|--C. clima Forster 1979PVD10
|--C. compta Koch 1839K55
|--C. consensa Forster 1979PVD10
|--C. contrita Forster 1979PVD10
|--C. convoluta Forster 1979PVD10
|--C. corrugata Bösenberg & Strand 1906 [incl. C. coerulescens sinensis Hu 1979]CH12
|--C. corticalisCH12
|--C. cycladataE07
|--C. damirkovaci Deeleman-Reinhold 2001CH12
|--C. deletrix P.-Cambridge 1885 [incl. C. maculata Song & Chen 1979, C. reichlini Schenkel 1944]CH12
|--C. delicata Forster 1979PVD10
|--C. diversa P.-Cambridge 1862CH12
|--C. elaphinesB96
|--C. epimelas [=Aranea epimelas]C01
|--C. erratica Koch 1836S06 [=Aranea erraticaC01, Cheiracanthium erraticumC01]
|--C. exilipes Lucas 1846E12
|--C. expansa Huang & Chen in Chen & Huang 2012CH12
|--C. femorocalcarata Huang & Chen in Chen & Huang 2012CH12
|--C. frigidula Thorell 1875S06
|--C. frisia Wunderlich & Schuett 1995BBM02
|--C. genevensis Koch 1866PO08
|--C. giulianetti Rainbow 1898M72
|--C. glauca Risso 1826R26
|--C. grisea Koch 1866S06
|--C. hilaris Simon 1878K55
|--C. huttoni Forster 1979PVD10
|--C. hystrixCH12
|--C. insulana Ono 1989CH12
|--C. japonica Koch 1878CH12
|--C. japonicola Bösenberg & Strand 1906 [incl. C. parajaponicola Schenkel 1963]CH12
|--C. jucunda (Karsch 1879) [=Liocranum jucundum; incl. C. mantis Bösenberg & Strand 1906, C. sakatensis Saito 1939]CH12
|--C. kayashimai Ono 1994CH12
|--C. kigabensis Strand 1915J98
|--C. kuanshanensis Ono 1994CH12
|--C. kurilensis Bösenberg & Strand 1906CH12
|--C. kurosawai Ono 1986CH12
|--C. labradorensis Marx 1891S06
|--C. laudabilis Simon 1909M90
|--C. lutescens Westring 1851BBM02
|--C. mandibularis Lucas 1846E12
|--C. maritimaR14
|--C. moesta Banks 1896PO08
|--C. moralis Song & Zhu in Song et al. 1991CH12
|--C. neglectaV09
|--C. norvegica Strand 1900S06
|--C. oblonga Lucas 1846E12
|--C. ornata Lucas 1846E12
|--C. pacifica Banks 1896S06
|--C. pallipes Lucas 1846E12
|--C. parvula Lucas 1846E12
|--C. peculiaris Koch 1873PVD10
|--C. pelasgicaK18
|--C. phragmites Koch 1843BBM02
|--C. picta Kulcz. 1885S06
|--C. producta Forster 1979PVD10
|--C. pteronetoidesCH12
|--C. reclusa Cambridge 1863BBM02
|--C. robustaM90
|--C. rufipes Lucas 1846E12
|--C. sapporensis Hayashi 1986CH12
|--C. scatula Forster 1979PVD10
|--C. subnotabilis Strand 1907J98
|--C. subrostrata Zhang & Hu 1991CH12
|--C. subtilis Koch 1867BBM02
|--C. taiwanica Ono 1994 [incl. C. bonicula Ono 1994]CH12
|--C. tanikawai Ono 1989CH12
|--C. terrestris Westring 1851SL14
|--C. torta Forster 1979PVD10
|--C. trivialis Koch 1843BBM02
|--C. ungavensis Marx 1891S06
|--C. viridis Risso 1826R26
|--C. viridula Ono 1989CH12
|--C. wolongica Zhu & An 1999CH12
|--C. yaginumai Hayashi 1989CH12
|--C. yangmingensis Hayashi & Yoshida 1993CH12
|--C. yoshidai Hayashi 1989CH12
`--C. zillaCH12

*Type species of generic name indicated


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[CH12] Chen, S.-H., & W.-J. Huang. 2012. The Spider Fauna of Taiwan—Araneae: Miturgidae, Anyphaenidae, Clubionidae. National Taiwan Normal University.

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[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[J98] Jäger, P. 1998. Das Typenmaterial der Spinnentiere (Arachnida: Acari, Amblypygi, Araneae, Opiliones, Pseudoscorpiones, Scorpiones, Uropygi) aus dem Museum Wiesbaden. Jahrbuecher des Nassauischen Vereins fuer Naturkunde 119: 81–91.

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[K55] Kraus, O. 1955. Spinnen von Korsika, Sardinien und Elba (Arach., Araneae). Senckenbergiana Biologica 36: 371–394.

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[M72] Monroe, R. 1972. Chelicerate type-specimens in the Queensland Museum. Memoirs of the Queensland Museum 16 (2): 291–307.

[PVD10] Paquin, P., C. J. Vink & N. Dupérré. 2010. Spiders of New Zealand: annotated family key and species list. Manaaki Whenua Press: Lincoln (New Zealand).

[PS08] Pinzón, J., & J. Spence. 2008. Performance of two arboreal pitfall trap designs in sampling cursorial spiders from tree trunks. Journal of Arachnology 36 (2): 280–286.

[PO08] Pluess, T., I. Opatovsky, E. Gavish-Regev, Y. Lubin & M. H. Schmidt. 2008. Spiders in wheat fields and semi-desert in the Negev (Israel). Journal of Arachnology 36 (2): 368–373.

[R14] Ramírez, M. J. 2014. The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History 390: 1–374.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

[SL14] Samu, F., G. Lengyel, É. Szita, A. Bidló & P. Ódor. 2014. The effect of forest stand characteristics on spider diversity and species composition in deciduous-coniferous mixed forests. Journal of Arachnology 42 (2): 135–141.

[S06] Strand, E. 1906. Die arktischen Araneae, Opiliones und Chernetes. In: Römer, F., & F. Schaudinn (eds) Fauna Arctica. Eine Zusammenstellun der arktischen Tierformen, mit besonder Berücksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nördliche Eismeer im Jahre 1898 vol. 4 pp. 431–478. Gustav Fischer: Jena.

[V09] Verdcourt, B. (ed.) 2009. Additions to the wild fauna and flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.

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