
Belongs within: Medusozoa.
The Codonophora is a clade within the siphonophores characterised by the presence of nectosomes, bell-shaped zooids specialised for swimming, and gelatinous shield-like bracts (these latter lost in some subgroups) (Dunn et al. 2005).
<==Codonophora [Brachystelia, Physonectae] | i. s.: Lychnagalma utricularia [incl. L. vesicularia]DPH05 | Discolabe quadrigataH04 | Strobalia cupolaH04 |--ApolemaDPH05 [ApolemidaeDB07] `--+--Cordagalma cordiformeDPH05 |--ErennaDPH05 |--Stephalia [Rhodaliidae]DPH05 | `--S. dilataDPH05 |--StephanomiaDPH05 | |--S. amphytridis [=Halistemma amphytridis, S. amphitridis (l. c.)]DPH05 | |--S. bijugaB26 | |--S. caraB26 | `--S. uvariaR26 |--PyrostephidaeDB07 | |--Pyrostephos vanhoeffeniPO99 | `--BargmanniaDPH05 | |--B. amoenaDPH05 | `--B. elongataDPH05 |--+--+--Physophora [Physophoridae]DPH05 | | | |--P. borealisM06 | | | |--P. hydrostaticaDPH05 | | | `--P. musonemaG20 | | `--Forskalia [Forskaliidae]DPH05 | | |--F. tholoidesDPH05 | | `--+--F. edwardsiDPH05 | | `--+--F. asymmetricaDPH05 | | `--F. formosaDPH05 | `--AgalmatidaeDPH05 | |--NanomiaDPH05 | | |--N. bijugaDPH05 | | `--N. caraIF02 | `--+--Halistemma rubrumDPH05 | `--AgalmaDPH05 | |--A. clausiDPH05 | `--+--A. elegansDPH05 | `--+--A. okeniDPH05 | `--Athorybia [Athorybiidae]DPH05 | `--A. rosaceaDPH05 `--Calycophorae [Calyconectae]DPH05 |--+--Praya [Prayidae]DPH05 | | |--P. dubiaDPH05 | | `--P. galeaH04 | `--+--Nectadamas diomedeaeDPH05 | `--Nectopyramis natansDPH05 `--+--Craseoa latheticaDPH05 |--+--+--Gymnopraia lapislazulaDPH05 | | `--Rosacea flaccidaDPH05 | `--Vogtia Kölliker 1853DPH05 | |--V. pentacanthaDPH05 | `--+--V. glabraDPH05 | `--Hippopodius Quoy & Gaimard 1827 [Hippopodiidae]DPH05 | `--H. hippopusDPH05 `--+--+--Chuniphyes multidentataDPH05 | `--Clausophyes [Clausophyidae]DPH05 | `--C. ovataDPH05 `--+--Sphaeronectes [Sphaeronectidae]DPH05 | |--S. fragilisPO99 | |--S. gracilisDPH05 | `--S. irregularisPO99 `--DiphyidaeDPH05 |--+--Muggiaea atlanticaDPH05 | `--+--Lensia conoideaDPH05 | `--Sulculeolaria quadrivalvisDPH05 `--+--Chelophyes appendiculataDPH05 `--+--AbylidaeDPH05 | |--Abylopsis tetragonaDPH05 | `--Bassia obeliscusH04 `--DiphyesDPH05 |--D. appendiculataB26 |--D. arcticaB26 |--D. bojaniB26 |--D. chamissonisZS10 |--D. disparDPH05 |--D. spiralisB26 `--D. truncataB26
*Type species of generic name indicated
References
[B26] Bigelow, H. B. 1926. Plankton of the offshore waters of the Gulf of Maine. Bulletin of the Bureau of Fisheries 40 (2): 1–509.
[DB07] Daly, M., M. R. Brugler, P. Cartwright, A. G. Collins, M. N. Dawson, D. G. Fautin, S. C. France, C. S. McFadden, D. M. Opresko, E. Rodriguez, S. L. Romano & J. L. Stake. 2007. The phylum Cnidaria: a review of phylogenetic patterns and diversity 300 years after Linnaeus. Zootaxa 1668: 127–182.
[DPH05] Dunn, C. W., P. R. Pugh & S. H. D. Haddock. 2005. Molecular phylogenetics of the Siphonophora (Cnidaria), with implications for the evolution of functional specialization. Systematic Biology 54 (6): 916–935.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.
[IF02] Ivantsov, A. Yu., & M. A. Fedonkin. 2002. Conulariid-like fossil from the Vendian of Russia: a metazoan clade across the Proterozoic/Palaeozoic boundary. Palaeontology 45 (6): 1219–1229.
[M06] Maas, O. 1906. Die arktischen Medusen (ausschliesslich der Polypomedusen). In: Römer, F., & F. Schaudinn (eds) Fauna Arctica. Eine Zusammenstellun der arktischen Tierformen, mit besonder Berücksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nördliche Eismeer im Jahre 1898 vol. 4 pp. 479–526. Gustav Fischer: Jena.
[PO99] Pagès, F., & C. Orejas. 1999. Medusae, siphonophores and ctenophores of the Magellan region. Scientia Marina 63 (Suppl. 1): 51–57.
[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.
[ZS10] Zhang, G.-T., S. Sun, Z.-L. Xu & Q.-L. Zhang. 2010. Unexpected dominance of the subtropical copepod Temora turbinata in the temperate Changjiang River estuary and its possible causes. In: China-Russia Bilateral Symposium: Proceedings of the China-Russia Bilateral Symposium of “Comparison on Marine Biodiversity in the Northwest Pacific Ocean”, 10–11 October 2010, Qingdao (China) pp. 100–102. Institute of Oceanology, Chinese Academy of Sciences; A. V. Zhirmunsky Institute of Marine Biology, Far East Branch of the Russian Academy of Sciences.