Coelocybidae

 Ormyromorpha sp., photographed by John M. Heraty.

Belongs within: Chalcidoidea.

The Coelocybidae are a group of small wasps whose larvae, so far as is known, develop in the galls created by other insects on various trees (Bouček 1988).

Characters (from Bouček 1988, as Coelocybinae): Inner eye margins diverging in lower or anterior section. Antennae often relatively short, with less sclerotised area of sensilla on enlarged clava, the segments of which are often fused. Pronotum rather short. Notauli obliterated. Scutellum with a pair of bristles on sides of the frenal line. Propodeum frequently with a basal ‘cup’.

<==Coelocybidae [Coelocybinae, Lieparini]BM22
    |--LanthanomyiaHB13
    |--Liepara Bouček 1988BM22, D07
    |    |--*L. dentata Bouček 1988B88
    |    `--L. dahmsi Bouček 1988B88
    |--Eucoelocybomyia Girault 1915B88
    |    `--*E. aerea Girault 1915B88
    |--Tomicobomorphella Girault 1915B88
    |    `--*T. justicia Girault 1915B88
    |--Nerotolepsia Girault 1920B88
    |    `--*N. bella Girault 1920B88
    |--Paratomicobia Girault 1915B88
    |    `--*P. flavios Girault 1915 [incl. Ormyromorphella bioculata Girault 1925, Tomicobomorphella heinei Girault 1922]B88
    |--Cooloolana Bouček 1988B88
    |    `--*C. albicoxa Bouček 1988B88
    |--Acoelocyba Bouček 1988B88
    |    `--*A. nemorum (Girault 1927) [=Erotolepsiella nemorum]B88
    |--Erotolepsiella Girault 1915B88
    |    `--*E. bifasciata Girault 1915B88
    |--Fusiterga Bouček 1988B88
    |    |--*F. gallarum Bouček 1988B88
    |    `--F. lativentris Bouček 1988B88
    |--Coelocyboides Girault 1913B88
    |    |--*C. pax Girault 1913B88
    |    `--C. bialbiguttus Girault 1933B88
    |--Cybopella Bouček 1988B88
    |    |--*C. eucalypti Bouček 1988B88
    |    `--C. tasmanica Bouček 1988B88
    |--Lelapsomorpha Girault 1913B88
    |    |--*L. myersi Girault 1913B88
    |    `--L. bioculata (Girault 1923) [=Coelocyboides bioculata]B88
    |--Ormyromorpha Girault 1913 [incl. Ormyromorphella Girault 1915]B88
    |    |--*O. trifasciatipennis Girault 1913B88
    |    |--O. aeneiscapus Girault 1927B88
    |    |--O. biargentinotata (Girault 1915) [=*Ormyromorphella biargentinotata]B88
    |    |--O. ciliata Girault 1925 [=O. glabra ciliata]B88
    |    |--O. glabra Girault 1915B88
    |    |--O. petiolata Girault 1925B88
    |    |--O. sexsetosa Girault 1925B88
    |    |--O. silvifilia Girault 1927B88
    |    |--O. thera (Walker 1839) [=Miscogaster thera, Lamprotatus thera]B88
    |    `--O. trifasciata Girault 1913B88
    |--Coelocyba Ashmead 1900 (see below for synonymy)B88
    |    |--*C. nigrocincta Ashmead 1900B88 (see below for synonymy)
    |    |--C. acincta Girault 1913 [=*Paracoelocyba acincta, *Pareunotus acinctus]B88
    |    |--C. eucalypti Girault 1934B88
    |    |--C. flavicorpus (Girault 1915) [=*Pareunotus flavicorpus]B88
    |    |--C. minuta (Girault 1916) (see below for synonymy)B88
    |    |--C. nigriventris (Girault 1915) [=Coelocybomyia nigriventris]B88
    |    |--C. persimilis (Girault 1915)GM79
    |    |--C. spenceri (Girault 1915) [=*Taneostigmomyia spenceri]B88
    |    |--C. turneri Girault 1931B88
    |    `--C. varifasciata Girault 1915B88
    |--AmbogasterBM22
    `--AriasinaBM22

Coelocyba Ashmead 1900 [incl. Coelocybomyia Girault 1915, Paracoelocyba Girault 1916, Pareunotus Girault 1915, Taneostigmomyia Girault 1915]B88

Coelocyba minuta (Girault 1916) [=Paracoelocyba minuta; incl. C. minuta iridos Girault 1931, P. minuta var. nigriventris Girault 1916 non Coelocybomyia nigriventris Girault 1915]B88

*Coelocyba nigrocincta Ashmead 1900B88 [=C. nigricincta (l. c.)B88; incl. Coelocybomyia novisexfasciata Girault 1915B88, Coelocybomyia persimilis Girault 1915B88, Coelocyba persimilisG34, *Coelocybomyia sexfasciata Girault 1915B88]

*Type species of generic name indicated

References

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[BM22] Burks, R., M.-D. Mitroiu, L. Fusu, J. M. Heraty, P. Janšta, S. Heydon, N. D.-S. Papilloud, R. S. Peters, E. V. Tselikh, J. B. Woolley, S. van Noort, H. Baur, A. Cruaud, C. Darling, M. Haas, P. Hanson, L. Krogmann & J.-Y. Rasplus. 2022. From hell’s heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera). Journal of Hymenoptera Research 94: 13–88.

[D07] Desjardins, C. A. 2007. Phylogenetics and classification of the world genera of Diparinae (Hymenoptera: Pteromalidae). Zootaxa 1647: 1–88.

[G34] Girault, A. A. 1934. New Capsidae and Hymenoptera, with note on an unmentionable. Privately published (reprinted: Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 311–314).

[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466–542.

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