Parvanachis obesa, copyright Muséum national d’histoire naturelle.

Belongs within: Nassariidae.
Contains: Liratilia, Zemitrella, Columbella, Pyrene, Costoanachis, Anachis, Mitrella, Amphissa, Aesopus, Astyris, Strombina, Zafra.

The Columbellidae, dove shells, are a group of small to medium-sized, active marine gastropods including both carnivorous and herbivorous species. The family is divided between two subfamilies: Pyreninae tend to have narrow fusiform shells, and have a radula with sigmoid lateral teeth and a narrow median tooth, whereas Columbellinae tend to have coniform or strombiform shells, and have a radula with broad and flat lateral teeth and a broader median tooth (Radwin 1977).

Within the Pyreninae, Parvanachis is a genus of small dove shells with a very squat shell ornamented with strong axial ribs (Radwin 1977). Conella species have medium-sized biconic shells that are smooth or bear microscopic spiral striae (Radwin 1978).

Characters (from Seashells of NSW): Siphonal canal very short, often little more than a gap at anterior end of aperture. Top of outer lip with sinus present, usually shallow; denticles usually present inside outer lip, occasionally just a single indistinct bulge. Inner (columellar) lip without plaits, weak denticles often present, aligned on an axial ridge just inside edge of callus.

<==Columbellidae [Atypoglossa, Columbelloidea, Pyrenidae, Pyrenoidea]
    |    |--ColumbellaR78
    |    `--Zafrona Iredale 1916 [incl. Metanachis Thiele 1924]W94
    |         |--*Z. isomella (Duclos 1840)W94, BD12 [=Columbella isomellaW94]
    |         |--Z. acleonta (Duclos 1840) (see below for synonymy)W94
    |         |--Z. lifouana (Hervier 1899)WBS-S15
    |         `--Z. pulchella (Blainville 1829) (see below for synonymy)BC01
    `--Pyreninae [Atiliidae, Atiliinae, Atilinae]R78
         |--Parametaria dupontii (Kiener 1849–1850)R78, PP78
         |--Eurypyrene Woodring 1928R78
         |--Rhombinella laevigata (Linnaeus 1758)KF22
         |--Atilia Adams & Adams 1853R78
         |    `--*A. suffusa (Sowerby 1844) [=Columbella suffusa]BR17
         |--Decipifus Olsson & McGinty 1958R78
         |    `--*D. sixaolus Olsson & McGinty 1958 [incl. Buccinum pulchellum Adams 1851 non Blainville 1829]R78
         |--Exomilopsis Powell 1964W94
         |    `--E. spica (Hedley 1907)P66 [=Mangilia spicaW94, Exomilus spicaP66]
         |--Gatliffena Iredale 1929W94
         |    `--*G. fenestrata (Tate & May 1900)W94 [=Donovania fenestrataW94, Exomilus fenestratusH22, Trophon fenestratusH22]
         |--Zella Iredale 1924W94
         |    `--*Z. beddomei (Petterd 1884) [=Terebra beddomei; incl. Z. attenuata Angas 1871]W94
         |--Mazatlania Dall 1900 [=Euryta Adams & Adams 1853 non Gistel 1848]R78
         |    |--*M. aciculata (Lamarck 1822)R78 (see below for synonymy)
         |    `--M. fulgurata (Philippi 1846)R78 (see below for synonymy)
         |--Pseudanachis Thiele 1924W94
         |    |--*P. duclosianus (Sowerby 1844)W94 [=Columbella duclosianaW94, Pyrene duclosianaH09]
         |    `--P. basedowi (Hedley 1918)KF22
         |--Columbellopsis Bucquoy & Dautzenberg in Bucquoy, Dautzenberg & Dollfus 1882R78
         |    |--*C. minor (Scacchi 1836) [=Columbella minor, Atilia minor, Mitrella minor]R78
         |    |--C. edmondi (Briart & Cornet 1870) [=Mitrella (Columbellopsis) edmondi]TTE93
         |    `--C. nycteis (Duclos in Chenu 1846) (see below for synonymy)R78
         |--Steironepion Pilsbry & Lowe 1932 [incl. Psarostola Rehder 1943]R77
         |    |--S. tincta (Carpenter 1864) (see below for synonymy)R77
         |    |--S. minor (Adams 1845) (see below for synonymy)R77
         |    `--S. monilifera (Sowerby 1844) (see below for synonymy)R77
         |--Suturoglypta Radwin 1968R78
         |    |--*S. pretri (Duclos in Chenu 1846) (see below for synonymy)R78
         |    |--S. albella (Adams 1850) [=Pleurotoma albella]R78
         |    `--S. iontha (Ravenel 1861) [=Columbella iontha; incl. Anachis acuta Stearns 1873]R78
         |--Cosmioconcha Dall 1913R78
         |    |--*C. modesta (Powys 1835)R78 (see below for synonymy)
         |    |--C. calliglypta (Dall & Simpson 1901) [=Anachis calliglypta]R78
         |    `--C. nitens (Adams 1850) (see below for synonymy)R78
         |--Parviterebra Pilsbry 1904W94
         |    |--*P. paucivolvis Pilsbry 1904W94
         |    |--P. brazieri (Angas 1875) (see below for synonymy)W94
         |    `--P. trilineata Adams & Angas 1863W94
         |--Pseudamycla Pace 1902W94
         |    |--*P. dermestoidea (Lamarck 1822) [=Buccinum dermestoidea; incl. P. lineolata Tryon 1883]W94
         |    |--P. formosaZLK11
         |    `--P. miltostoma (Tenison Woods 1876)W94
         |--Metulella Gabb 1873R77
         |    |--*M. fusiformis Gabb 1873R77
         |    |--M. columbellata Dall 1889 [=M. (Nassarina) columbellata]R77
         |    |--M. venustaR77
         |    `--M. williamgabbiR77
         |--Zanassarina Pilsbry & Lowe 1932R77
         |    |--*Z. poecilaR77
         |    |--Z. albipesR77
         |    |--Z. atellaR77
         |    `--Z. xenoR77
         |--Graphicomassa Iredale 1929W94
         |    |--*G. ligula (Duclos 1840) (see below for synonymy)W94
         |    |--G. adiostina (Duclos 1840)KF22
         |    |--G. albina (Kiener 1841)WBS-S15
         |    |--G. puella (Sowerby 1844) (see below for synonymy)W94
         |    `--G. venulata (Sowerby 1894) [=Mitrella (Graphicomassa) venulata]W94
         |--Retizafra Hedley 1913W94
         |    |--*R. gemmulifera (Hedley 1907)F27, H09 [=Pyrene gemmuliferaF27]
         |    |--R. brevissima Hervier 1899W94
         |    |--R. calva (Verco 1910)W94
         |    |--R. intricata (Hedley 1912)W94 [=Columbella clathrata Brazier 1877W94, Pyrene clathrataH09]
         |    `--R. plexa (Hedley 1902)W94
         |--Nassarina Dall 1889R77
         |    |  i. s.: N. rietae Segers & Swinnen 2004F05
         |    |--N. (Nassarina)R77
         |    |    |--*N. (N.) bushii (Dall 1889) [=Nassaria (*N.) bushii]R77
         |    |    |--N. (N.) glypta (Bush 1885) (see below for synonymy)R77
         |    |    `--N. (N.) metabrunnea Dall & Simpson 1901R77
         |    `--N. (Cigclirina) helenae Keen 1971BC01
         |--Euplica Dall 1889W94
         |    |--*E. turturina (Lamarck 1822)W94, BW09
         |    |--E. deshayesii (Crosse 1859)BW09
         |    |--E. festiva (Deshayes 1834)BD12
         |    |--E. ionida (Duclos 1840)BD12
         |    |--E. livescens (Reeve 1859)WBS-S15
         |    |--E. scripta (Lamarck 1822)WBS-S15
         |    |--E. turturina (Lamarck 1822)WBS-S15
         |    `--E. varians (Sowerby 1832)BD12 [incl. Pyrene (Columbella) nodulosa Usticke 1959BC01]
         |--Conella Swainson 1840 [=Cionella (l. c.) non Jeffreys 1830]R78
         |    |--C. ovulata (Lamarck 1822) [=Columbella ovulata, Pyrene ovulata, *Conella picata Swainson 1840]R78
         |    |--C. cedonulli [=Meta cedonulli]C64
         |    |--C. coniformis [=Meta coniformis]C64
         |    |--‘Meta’ dupontiaeC64
         |    |--C. ledaluciae (Rios & Tostes 1981) [=Pyrene (Conella) ledaluciae]BC01
         |    |--C. ovuloides (Adams 1850) (see below for synonymy)R78
         |    `--C. perplexabilis (Maury 1917) [=Meta perplexabilis]R78
         |--Macrozafra Finlay 1927W94
         |    |--*M. subabnormis (Suter 1899) [=Clathurella subabnormis]P61
         |    |    |--M. s. subabnormisP61
         |    |    `--M. s. saxatilis (Murdoch 1905)P61 [=Columbella saxatilisF27]
         |    |--M. calva [=Pyrene calva]F27
         |    |--M. cophinodes (Suter 1908)P61
         |    |--M. enwrighti Powell 1940P61
         |    |--M. mariae Powell 1940P61
         |    `--M. nodicincta (Suter 1899)P61 [=Clathurella nodicinctaF27]
         `--Parvanachis Radwin 1968R77
              |--*P. obesa (Adams 1845) (see below for synonymy)R77
              |--P. decorata (Strebel 1905)R78
              |    |--P. d. decorataR78
              |    `--P. d. inornataR78
              |--P. isabellei (Orbigny 1839) (see below for synonymy)R77
              |--P. melvillei (Strebel 1905)R78 [=Columbella (Seminella) melvilleiR77]
              |--P. ostreicola (Sowerby 1882) [=Columbella (Anachis) ostreicola, Anachis obesa ostreicola]R77
              |--P. paessleri (Strebel 1905)R78
              `--P. rhodae (Radwin 1968) [=Anachis (Parvanachis) rhodae]R77
Columbellidae incertae sedis:
  Antizafra Finlay 1927P61
    |--*A. pisaniopsis [=Columbella pisaniopsis]P61
    |--A. cancellaria [=Columbella cancellaria]F27
    |--A. plexa [=Columbella plexa]F27
    |--A. speighti [=Anachis speighti]F27
    `--A. vivens Powell 1934P61
  Paxula Finlay 1927P61
    |--*P. paxillus (Murdoch 1905) [=Columbella paxillus]P61
    |--P. allani Finlay 1928P61
    |--P. leptalea (Suter 1908)P61 [=Mitrella leptaleaF27]
    |--P. murdochi Finlay 1927P61
    |--P. sorenseni Powell 1955P61
    |--P. subantarctica (Suter 1908)P61 [=Mitrella subantarcticaF27]
    `--P. transitans (Murdoch 1905)P61 [=Columbella transitansF27]
  Pardalina testudinaria (Link 1807)BW09
  Nitidella Swainson 1840O27
    |--*N. nitida [=Columbella nitida]O27
    |--N. densilineataC64
    |--N. gouldi Carpenter 1856O27
    |--N. luculenta Dall 1919O27
    `--N. millepunctataC64
  Cotonopsis Olsson 1942BC01
    |  i. s.: C. monfilsi Emerson 1993BC01
    |         C. phuketensis (Kosuge, Roussy & Muangman 1998) (see below for synonymy)BC01
    |--C. (Cotonopsis)BC01
    |    |--C. (C.) deroyae (Emerson & D’Attilio 1969) [=Strombina (C.) deroyae]BC01
    |    |--C. (C.) mendozana (Shasky 1970) [=Strombina (C.) mendozana]BC01
    |    |--C. (C.) skoglundae Jung 1989BC01
    |    `--C. (C.) suteri Jung 1989BC01
    `--C. (Turrina) turrita [incl. Strombina elegans Li 1930]BC01
  Ruthia Shasky 1970BC01
    `--*R. mazatlanica Shasky 1970BC01
  Zetekia Dall 1918BK11, P66
    |--Z. gemmulosa (Adams 1852) [=Pleurotoma gemmulosa; incl. *Z. denticulata Dall 1918]P66
    `--Z. curta Dall 1920P66
  Antimitrella Powell 1937P61
    `--*A. laxa Powell 1937P61
  Aoteatilia Powell 1939P61
    |--*A. substriata (Suter 1899) [=Daphnella substriata]P61
    |--A. acicula (Suter 1908)P61
    |--A. amphipsila (Suter 1908)P61
    |--A. larochei Powell 1940P61
    |--A. psila (Suter 1908)P61
    `--A. tenuistriata (Suter 1908)P61
  *Seminella varia (Pease 1860)K65, H22, K65 [=Cythara variaH22]
  Mokumea fuscolineata (Thiele 1830)WBS-S15
  Pardalinops marmorata (Gray 1839)WBS-S15
  Pictocolumbella ocellata (Link 1807)WBS-S15

Columbellopsis nycteis (Duclos in Chenu 1846) [=Colombella nycteis; incl. Columbella belizana Duclos in Chenu 1848, Columbella fenestrata Adams 1850, Anachis (Alia) fenestrata, Mitrella (Columbellopsis) fenestrata, Colombella fusiformis d’Orbigny in Sagra 1842 non Anton 1839, Colo. plutonida Duclos in Chenu 1846]R78

Conella ovuloides (Adams 1850) [=Columbella ovuloides, Col. (Conoidea) ovuloides, Meta ovuloides, Pyrene ovuloides]R78

*Cosmioconcha modesta (Powys 1835)R78 [=Buccinum modestumR78, Amphissa (Cosmioconcha) modestaBC01; incl. Strombina laevistriata Li 1930BC01]

Cosmioconcha nitens (Adams 1850) [=Fusus nitens; incl. Columbella (Astyris) perpicta Dall & Simpson 1901, Mitrella perpicata (l. c.)]R78

Cotonopsis phuketensis (Kosuge, Roussy & Muangman 1998) [=Strombina phuketensis; incl. C. wanwalleghemi Kronenberg & Dekker 1998]BC01

*Graphicomassa ligula (Duclos 1840) [=Columbella ligula, Mitrella (*Graphicomassa) ligula; incl. M. carnea Hervier 1899, M. insignis Adams 1853, M. nivalis Hervier 1899]W94

Graphicomassa puella (Sowerby 1844) [=Mitrella (Graphicomassa) puella; incl. M. baculus Reeve 1859, M. brevis Schepman 1913, M. circulata Hervier 1899, M. conspersa Gaskoin 1852, M. contaminata Gaskoin 1852, M. diluta Hervier 1899, M. intemerata Hervier 1899, M. nodosa Schepman 1913, M. sigaloessa Melvill & Standen 1896, M. suspecta Hervier 1899]W94

*Mazatlania aciculata (Lamarck 1822)R78 [=Buccinum aciculatumR78, Euryta aciculataC64, Terebra aciculataH43; incl. T. consentini Philippi 1826R78, Mazatlania hesperia Pilsbry & Lowe 1932R78, T. nodosoplicata Dunker 1853R78]

Mazatlania fulgurata (Philippi 1846)R78 [=Terebra fulgurataR78, Euryta fulgurataC64; incl. T. arguta Gould 1853R78, Aesopus goforthi Dall 1912R78]

Nassarina (Nassarina) glypta (Bush 1885) [=Mangelia glypta, Nassaria (N.) glypta; incl. Nassarina dalli Olsson & Harbison 1953, Columbella (Atilia) mystica Dall 1927]R77

Parvanachis isabellei (Orbigny 1839) [=Buccinum isabellei, Anachis isabellei, Columbella isabellei, Nassa isebellei; incl. Anachis (Parvanachis) radwini Altena 1975]R77

*Parvanachis obesa (Adams 1845) [=Buccinum obesum, Anachis obesa, Costoanachis obesa; incl. Buccinum concinnum Adams 1845 non Dillwyn 1817, Columbella costulata Adams 1850 non Cantraine 1835, Col. crassilabris Reeve 1859, Col. decipiens Adams 1850, Col. ornata Ravenel 1859, Col. pygmaea Adams 1850]R77

Parviterebra brazieri (Angas 1875) [incl. P. angasi Tryon 1884, P. australis Tenison Woods 1878, P. harrisoni Tenison Woods 1877]W94

Steironepion minor (Adams 1845) [=Pleurotoma minor; incl. Colombella monilifera Duclos in Chenu 1846 non Columbella monilifera Sowerby 1844]R77

Steironepion monilifera (Sowerby 1844) [=Columbella monilifera; incl. Pleurotoma fuscolineata Adams 1845, Psarostola monilifera sparsipunctata Rehder 1943, Steironepion monilifera sparsipunctata, Columbella telea Duclos in Chenu 1848]R77

Steironepion tincta (Carpenter 1864)R77 [=Anachis tinctaR77, Nassarina (Steironepion) tinctaPP78; incl. Mangelia melanosticta Pilsbry & Lowe 1932R77, *Steironepion melanostictaR77]

*Suturoglypta pretri (Duclos in Chenu 1846) [=Colombella pretrii, Columbella pretrei; incl. Columbella mangelioides Reeve 1859, Anachis samanensis Dall 1889]R78

Zafrona acleonta (Duclos 1840) [incl. Columbella jaspidea Sowerby 1844, *Metanachis jaspidea, Zafrona plicaria Montrouzier 1862]W94

Zafrona pulchella (Blainville 1829) [incl. Anachis multicostata Usticke 1969, A. subcostulata var. subcincta Usticke 1969]BC01

*Type species of generic name indicated


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