Belongs within: Curculioninae.
Contains: Lechriopini, Zygopini, Peridinetus, Ambates, Tonesiina, Nertinina, Torcina, Coelonertina, Diorymerina, Madarina, Lydamis, Eurhinina, Madopterina, Pseudobaris, Heterosternus, Baris, Solenosternus, Anopsilus.
The Conoderinae are a diverse group of weevils united by the presence of a strongly curved submarginal fold on the ventrolateral surface of the elytra (Davis 2009). The monophyly of this group is uncertain and its major subgroups may be treated as distinct subfamilies. These include the the Conoderitae, in which the eyes are large and usually subcontiguous, separated dorsally by only a very narrow forehead (Legalov 2018). The Orobitidini have a spherical body form, with ventrites 1 to 4 subequal in length, and an exposed pygidium (Legalov 2018).
The Bariditae are a large group of about 550 genera with a more elongate body form (Legalov 2018). The morphological diversity within the group is notably high, and the baridines are consequently divided between a number of tribes and subtribes.
<==Conoderinae [Campyloscelinae] | i. s.: Diomia tetragramma Pascoe 1885Mac86 | Illacuris laticollisMas86 | Cratosomus fasciatusML05 | TomicoproctusZ93 | Philinna Champion 1906A02 | Philides Champion 1906A02 |--Orobitidini [Orobitiditae, Orobitidinae]L18 | |--ParorobitisD09 | | |--P. gibbus Korotyaev, O’Brien & Konstantinov 2000D09 | | `--P. minutusB14 | `--Orobitis Germar 1817L18 | |--O. cyaneaB14 | `--O. nigrinaB14 |--ConoderitaeL18 | |--CorynemeriniL18 | |--PiazurusLB20 [PiazuriniL18] | |--PeloropodiniL18 | |--LechriopiniL18 | |--ZygopiniL18 | |--Trichodocerus Chevrolat 1879D09 [TrichodoceriniL18] | | `--T. spinolae Chevrolat 1879L18 | |--CyllophorusD09 [CoryssopodiniL18, Sympiezopides] | | `--C. fasciatus Faust 1886D09 | |--OthippiaMas86 [OthippiiniL18] | | `--O. guttula Pascoe 1885Mas86 | |--Conoderes [Conoderini]L18 | | `--C. albidus Schoenherr 1833L18 | |--SphadasmusHF03 [SphadasminiL18] | | |--S. camelus (Gyllenhal 1836)L18 | | `--S. sinaiticus Pic 1919HF03 | |--Metialma Pascoe 1871D09, M94 [CoryssomeriniL18, Metialmini, Synophthalmides] | | |--M. anisomelisM94 | | |--M. balsaminaeM94 | | `--M. signifera Pascoe 1871D09 | |--Lobotrachelus Schoenherr 1838M94 [LobotracheliniL18] | | |--L. exilis Pascoe 1885Mas86 | | |--L. stigma Pascoe 1873Mas86 | | `--L. troglodytes (Desbrochers 1891)D09 | `--Menemachini [Menemachides]L18 | |--Balanogastris kolae (Desbrochers 1895)D09 | |--Telephae oculata (Say 1824)D09 | `--Berethia sannio Pascoe 1872Mas86 `--Bariditae [Barididae, Baridides, Baridinae, Barinae, Madopterini, Centrinini]L18 | i. s.: Pseudocholus cinctusMac86 | ‘Curculio’ curvirostris Fabricius 1781Z93 | Angelocentris schubarti Bondar 1950 [incl. Leptoschoinella carinirostris Hustache 1951]K83 | Rhaptinus Casey 1920K83 | `--R. tabaci [=Baris tabaci]K83 | Bonomius Casey 1922K83 [incl. Eurhinopsella Hustache 1950 K83, Eurhonopsellix (l. c.)H49] | `--B. aeneoviridis Casey 1922K83 [incl. *Eurhinopsella triangularis Hustache 1950H49, K83] | Castenus pyritosus (Germar 1824) [incl. C. cupreolatus Casey 1922]K83 | Eurypages pennatus Pascoe 1872 [incl. E. rectirostris Casey 1922]K83 | HulpesellusK83 | |--H. delicatulus [=Myelantiella delicatula]K83 | `--H. vitraci [=Notesia vitraci]K83 | Lispodemus germari (Boheman 1844) [=Nertinus germari; incl. L. femoralis Lacordaire 1866]K83 | ParamadarusK83 | |--P. complexus Casey 1922 [incl. P. bruchi Hustache 1926]K83 | |--P. gounellei (Hustache 1951)K83, H51b [=Linomadarus gounelleiK83] | |--P. migrator [=Linomadarus migrator]K83 | `--P. similis (Hustache 1951)K83, H51b [=Linomadarus similisK83] | Paratorcus Casey 1922 [incl. Atorcus Bondar 1943]K83 | `--P. angulicollis [=Limnobaris angulicollis]K83 | Physoproctus Lacordaire 1866 [incl. Attangus Casey 1922, Madaropsides Casey 1922]K83 | |--‘Madaropsides’ biimpressus Hustache 1951H51b | `--P. glabratus (Gyllenhal 1836) (see below for synonymy)K83 | Stethobaropsis Casey 1922 [incl. Dahlgrenia Bondar 1942]K83 | Theantis striatula Casey 1922 [incl. T. pectoralis Casey 1922]K83 | Zalinus cribriventris (Hustache 1951)K83, H51b [=Madarus (Eumadarus) cribriventrisK83, H51b] | Barinomorphus Morimoto 1962M62 | `--*B. antennatus Morimoto 1962M62 | Eremonychus Marshall 1935M62 | Pseudorhyssematus Morimoto 1962M62 | `--*P. rufitarsis (Roelofs 1875) [=Rhyssematus rufitarsis]M62 |--PeridinetusD09 |--Ambatini [Pteracanthides]D09 | |--AmbatesD09 | `--Pteracanthus smidtii Dejean 1835D09 |--EutoxiniL18 | |--TonesiinaL18 | `--Eutoxus Schoenherr 1844 [Eutoxina]D09 | `--E. laeviusculus Hustache 1951K83, H51b [=E. laevisculusK83] |--Pantotelini [Pantotelides]D09 | |--Liturgus irrasusMas86 | `--Pantoteles Schönh. 1845 [Pantotelina]FS90 | |--P. subrectirostris Hustache 1949H49 | |--P. tenuirostris Boheman 1845D09 | `--P. variabilis Chv. 1879FS90 |--ApostasimeriniL18 | |--NertininaL18 | |--TorcinaL18 | |--CoelonertinaL18 | |--DiorymerinaL18 | `--Apostasimerus [Apostasimerina]L18 | `--A. serrirostris Boheman 1844L18 |--MadariniD09 | |--MadarinaD09 | |--NeosharpiinaL18 | | |--Dendrobaris Egorov 1976L18 | | |--Pharcidobaris Morimoto & Yoshihara 1996L18 | | |--Pellobaris Morimoto & Yoshihara 1996L18 | | |--Moreobaris Morimnoto & Yoshihara 1996L18 | | | `--M. deplanata (Roelofs 1875)D09 | | `--Orchidophilus Buchanan 1935L18 | | |--*O. peregrinator Buchanan 1935M94 | | `--O. aterrimus (Waterhouse 1874)D09 [=Acythopeus aterrimusM94] | `--Barymerina [Barymerini]D09 | |--PhysotroctusH51b | |--PhysoproetusH51b | |--Chapatiella Hustache 1951H51b | | `--*C. excelsa Hustache 1951H51b | |--Crassiopus Hustache 1951H51b | | `--*C. semicastaneus Hustache 1951H51b | `--Barymerus Lacordaire 1866 [incl. Myelantia Casey 1922]K83 | |--B. binarius Hustache 1938D09 | `--B. hispidulus [=Myelantia hispidula; incl. M. setosula Casey 1922]K83 |--Optatini [Cyrionychini]L18 | |--LydamisD09 | |--Optatus palmaris Champion 1907D09 | |--Telemus Pascoe 1889D09 | | |--T. chapadanus (Casey 1922) [incl. T. objectus Casey 1922]K83 | | `--T. dispersus Hustache 1949 [=T. griseopunctatus ms]H49 | |--PseudoptatusH49 | | |--P. dentipesH49 | | `--P. morio Hustache 1949H49 | |--Pistus Faust 1892D09, K83 [incl. Justus Bondar 1946K83, Neoscambus Hustache 1938K83, Omogonia Casey 1922K83] | | |--P. dromedarius (Kirsch 1875) [=Neoscambus dromedarius; incl. Justus gibber Bondar 1946, J. gibbifer]K83 | | |--P. galeatus (Boheman 1844)D09 (see below for synonymy) | | `--P. niveus (Hustache 1950) [=Neoscambus niveus; incl. Omogonia nivea Voss 1954]K83 | |--CostoviaH49 | | |--C. brunnea Hustache 1949H49 | | |--C. rubricollis Hustache 1949H49 | | `--C. tenuirostris Casey 1922 [incl. Eurypages tenuirostris Bondar 1946]K83 | |--PardisomusD09 | | |--P. biplagiatus (Desbrochers 1906)D09 | | |--P. guttatusH49 | | `--P. multiguttatus Hustache 1949H49 | |--AnonesH49 | | |--A. episternalis (Hustache 1949) [=Cyrionyx (Anones) episternalis]H49 | | |--A. fulvomaculatus (Hustache 1949) (see below for synonymy)H49 | | |--A. proximus [=Procholus proximus]K83 | | |--A. pulchellus [=Procholus pulchellus]K83 | | `--A. variegatus [=Conotrachelus variegatus]K83 | `--Cyrionyx [Cyrionychina]D09 | |--C. alboscutellatus Hustache 1949H49 | |--C. bruchi Hustache 1949H49 | |--C. camelus Champion 1907D09 | |--C. cinereus Hustache 1949H49 | |--C. maculatus [=Peridinetus maculatus]K83 | |--C. nitidulus Hustache 1949H49 | `--C. rufipes Hustache 1949H49 `--BaridiniD09 | i. s.: Melanobaris laticollisGC-P14 | HypogymniusK83 | Barinomorphoides Morimoto 1962M62 | `--*B. similaris Morimoto 1962M62 | Aulacobaris coeruescensHL23 | LabiaticolaHL23 |--EurhininaD09 |--Parallelodemas Faust 1894 [Parallelodemina]L18 |--MadopterinaL18 |--ColeomerinaD09 | |--Harotreus triangularis Hustache 1949H49 | `--ColeomerusD09 | |--C. abdominalis Hustache 1950L18 (see below for synonymy) | |--C. boliviensis Hustache 1949D09 | |--C. columbicus Hustache 1949H49 | `--C. subviolaceus Hustache 1949H49 `--Baridina [Anopsilini, Barina]D09 |--Genevra dentipes Hustache 1951H51a |--Sindesus ruber Hustache 1951 [=Prosindesus ruber ms]H51a |--GiveniusH51a |--EugiveniusH51a |--Anagivenius unidentatus Hustache 1951 [incl. A. unidentatus var. sanguinicollis Hustache 1951]H51a |--PseudobarisD09 |--HeterosternusH51a |--Sclerobaris violacea Hustache 1951H51a |--TrogatiaH51a |--Megops morosusH51a |--BarisD09 |--AcanthobarisH51a |--Craptus tibialis Casey 1922D09 |--Cryptosternum nevermanni Heller 1936D09 |--Dalcesia Casey 1922D09 |--Deipyle induta Champion 1909D09 |--Elasmobaris signifer Reitter 1895D09 |--Lepidobaris acnisti Champion 1909D09 |--Macrobaris Champion 1909D09 |--Megalobaris viridana (Boheman 1836)D09 |--Odontobaris Champion 1909D09 |--SolenosternusD09 |--Ulobaris loricata Reitter 1895D09 |--AnopsilusL18 |--Aulobaris LeConte 1876A02 |--Hesperobaris Casey 1892A02 |--Cosmobaris Casey 1920A02 | `--C. americana Casey 1920D09 |--Desmoglyptus Casey 1892A02 | `--D. arizonicus Casey 1920D09 |--Plesiobaris Casey 1892A02 | `--P. albilata (LeConte 1876)D09 |--Pycnobaris Casey 1892A02 | `--P. pruinosa Casey 1892D09 |--Rhoptobaris LeConte 1876A02 | `--R. canescens LeConte 1876D09 |--Trichobaris LeConte 1876A02 | `--T. texana LeConte 1876D09 |--Sphenobaris Champion 1909D09, K83 [incl. Amphibaris Casey 1922K83] | `--S. quadridens Champion 1909D09 |--Orthoris LeConte 1876L18 | `--O. crotchii LeConte 1876D09 |--Opertes Casey 1922D09 | `--O. glaber Hustache 1951H51a |--Mimobaris Hustache 1951H51a | `--*M. nitens Hustache 1951H51a |--Hasidellus Hustache 1951H51a | `--*H. robustus Hustache 1951 [=Hasidus robustus]H51a |--Claveriella Hustache 1951H51a | `--*C. violacea Hustache 1951H51a |--Plasiella Hustache 1951H51a | `--*P. subcylindrica Hustache 1951H51a |--Paranathaniops Hustache 1951H51a | `--*P. subfasciata Hustache 1951H51a |--Liogivenius Hustache 1951H51a | `--*L. ebeninus Hustache 1951H51a |--Buprestobaris Hustache 1951H51a | `--*B. caudata Hustache 1951H51a |--Scleronychius Hustache 1951H51a | `--*S. ebeninus Hustache 1951H51a |--Allomegops Hustache 1951D09 | `--*A. sulcatirostris Hustache 1951H51a |--Desmobaris Hustache 1951H51a | `--*D. singularis Hustache 1951H51a |--Trogatiella Hustache 1951H51a | `--*T. rufipes (Hustache 1924) [=Limnobaris rufipes]H51a |--Acanthobarisella Hustache 1951H51a | `--*A. antennata Hustache 1951H51a |--Microbaris Casey 1892A02 | `--M. galvestonica Casey 1892A02 |--Stenobaris Linell 1897A02 | `--S. avicenniae Linell 1897A02 |--Trepobaris Casey 1892A02 | |--T. elongata Casey 1892A02 | `--T. inornata Champion 1909D09 |--Nanobaris Champion 1909D09 [incl. Giveniopsis Casey 1922K83] | |--‘Giveniopsis’ nigrescens Hustache 1951 [=G. flavescens ms]H51a | `--‘Giveniopsis’ setosella Hustache 1951H51a |--BusckiellaD09 | |--B. lecythidis (Champion 1909)D09 | `--B. scutellata Hustache 1951H51a |--RhytidobarisH51a | |--R. seriesetosa Hustache 1951H51a | `--R. sulcata Hustache 1951H51a |--Archobaris Hustache 1951H51a | |--*A. undulata Hustache 1951H51a | `--A. brachypennis Hustache 1951H51a |--Coryssobaris Hustache 1951H51a | |--*C. sculpturata Hustache 1951 [=Baris lemoulti ms]H51a | `--C. malefida Hustache 1951 [=Baris malefida ms]H51a |--LupulinaH51a | |--L. brunnescens Hustache 1951H51a | `--L. subpubescens Hustache 1951H51a |--Anthobaris Hustache 1951H51a | |--*A. rugosicollis Hustache 1951H51a | `--A. denticulata Hustache 1951H51a |--Hispibaris Hustache 1951H51a | |--*H. albosetosa Hustache 1951H51a | `--H. unguicularis Hustache 1951K83, H51a [=H. onguicularisK83] |--DissopygusD09 | |--D. apicalis Hustache 1951H51a | |--D. catenulatus (Boheman 1836) [incl. D. oblongus Casey 1922]K83 | `--D. metallescens (Boheman 1836)D09 |--CymatobarisD09 | |--C. amethystina Hustache 1951H51a | |--C. coeruleaH51a | `--C. impressifrons Casey 1922D09 |--CyanobarisH51a | | i. s.: C. rufiventrisH51a | `--C. (Hedybaris Hustache 1951)H51a | `--C. (*H.) elegans Hustache 1951H51a |--Zathanius Casey 1922D09 [incl. Euzathanius Casey 1922K83] | |--Z. elongatulus Hustache 1951H51a | |--Z. lyratusH51a | `--Z. scutellatus Hustache 1951H51a |--EugymnobarisH51a | |--E. bosqi Hustache 1951 [=Gymnobaris bosqi ms]H51a | |--E. bruchi Hustache 1951 [=Gymnobaris bruchi ms]H51a | `--E. lazulina Hustache 1951 [=Gymnobaris lazulina ms]H51a |--CercobarisH51a | |--C. convexipennis Hustache 1951H51a | |--C. obtusata Hustache 1951H51a | `--C. rugosa Hustache 1951H51a |--OligobarisH51a | |--O. brevipilosus Hustache 1951H51a | |--O. cavicollis Hustache 1951H51a | `--O. nitidus Hustache 1951H51a |--LiobaridiaH51a | |--L. distinguenda Hustache 1951H51a | |--L. dubitabilisH51a | |--L. punctatella Hustache 1951K83, H51a | `--L. vicina Hustache 1951H51a |--RhytidoglymmaD09 | |--R. aenescens Faust 1896 [incl. Trachybaris willinki Bondar 1950]K83 | |--R. benoisti Hustache 1951H51a | |--R. caudata Hustache 1951H51a | |--R. sponsor Hustache 1951 [=Baris sponsor ms]H51a | `--R. subglabra Hustache 1951H51a |--MesothaniusH51a | |--*M. emarginatusH51a | |--M. canaliculatus Hustache 1951H51a | |--M. coelosternoides Hustache 1951K83, H51a [=M. coelosthernoidesK83] | |--M. sexpunctatus Hustache 1951 [=M. sexpustulatus ms]H51a | `--M. sparsus Hustache 1951H51a |--StereobarisD09 | |--S. albosetosa Hustache 1951 [=S. albosquamulata ms]H51a | |--S. impressithorax Hustache 1951H51a | |--S. interpunctata Casey 1922D09 | |--S. metallica (Boheman 1836) [incl. S. caseyi Hustache 1938, S. impressa Casey 1922]K83 | |--S. pardoensis Hustache 1951H51a | |--S. purpurea Hustache 1951H51a | `--S. revoili Hustache 1951H51a |--Methyorhina Pascoe 1872H51a, K83 | |--M. balteata (Casey 1922) [=Baris balteata; incl. Rhytidobaris brunneopilosa Hustache 1924]K83 | |--M. griseola Hustache 1951H51a | |--M. guyanea Hustache 1951H51a | |--M. hirsutipennis (Hustache 1951)K83, H51b [=Polpones hirsutipennisK83, Polponella hirsutipennis msH51b] | |--M. seriatosetulosa Hustache 1951H51a | |--M. squamulata Hustache 1951H51a | |--M. squamulosa [=Baris squamulosa]K83 | `--M. vestita Hustache 1951H51a |--Thanius Casey 1922D09 | |--T. columbicus Hustache 1951H51a | |--T. explanatus Hustache 1951H51a | |--T. impressirostris Hustache 1951H51a | |--T. perpolitusH51a | |--T. rufipes Hustache 1951H51a | |--T. sculpturatus Hustache 1951H51a | |--T. spadiceus (see below for synonymy)H51a | `--T. viridimicans Hustache 1951H51a `--BaptobarisH51a |--B. benoisti Hustache 1951H51a |--B. brevipennis Hustache 1951H51a |--B. claveri Hustache 1951H51a |--B. curta Hustache 1951H51a |--B. exarata [incl. B. e. var. semicastanea Hustache 1951]H51a |--B. gounellei Hustache 1951H51a |--B. rubripectus Hustache 1951K83, H51a [=B. rubropectusK83] `--B. ruficollis [incl. B. r. var. rufirostris Hustache 1951]H51a
Anones fulvomaculatus (Hustache 1949) [=Cyrionyx (Anones) fulvomaculatus, C. (Anonea [l. c.]) fulvomaculatus]H49
Coleomerus abdominalis Hustache 1950L18 [=C. rufipes msH49; incl. C. abdominalis var. vicinus Hustache 1949H49]
Physoproctus glabratus (Gyllenhal 1836) [=Attangus glabratus; incl. Madaropsides atrolucens Casey 1922, A. caudalis Casey 1922, Physoproctus dohrni Lacordaire 1866]K83
Pistus galeatus (Boheman 1844)D09 [=Scambus galeatusH49, *Neoscambus galeatusH49; incl. Omogonia cameloides Voss 1954K83, Justus tuberosus Bondar 1946K83]
Thanius spadiceus [incl. T. spadiceus var. basalis Hustache 1951, T. spadiceus var. bolivianus Hustache 1951, T. spadiceus var. insperatus Hustache 1951]H51a
*Type species of generic name indicated
References
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