Diagnostic figures of generalised Copidognathus, from Bartsch (2006).

Belongs within: Halacaridae.

Copidognathus is a large genus of primarily marine mites (though a few live in fresh water). Species are found living on a wide variety of substrates. They are distinguished by the absence of setae on the penultimate palp segment (Luxton 1990).

Characters (from Luxton 1990): Female with three pairs of perigenital setae and one pair of subgenital setae; palps four-segmented, penultimate segment lacking setae; genu I shorter than either femur I or tibia I; tibia I with three unpaired ventral setae.

<==Copidognathus Trouessart 1888 [incl. Copidognathopsis Viets 1927]N47
    |--C. (Copidognathus)N47
    |    |--*C. (C.) oculatus (Hodge 1863)L90 [=Halacarus oculatusT96, Copidognathopsis oculatusM62]
    |    |--C. (C.) acutus Newell 1947N47
    |    |--C. (C.) adriaticaN47
    |    |--C. (C.) angustus Viets 1936N47
    |    |    |--C. a. angustusN47
    |    |    `--C. a. floridensis Newell 1947N47
    |    |--C. (C.) bairdi Newell 1947N47
    |    |--C. (C.) biscayneus Newell 1947N47
    |    |--C. (C.) bonairensis (Viets 1936) [=Copidognathopsis bonairensis]N47
    |    |--C. (C.) bouvieri (Trouessart 1914) [=Werthella bouvieri]N47
    |    |--C. (C.) caudatus Newell 1947N47
    |    |--C. (C.) curassaviensis Viets 1936N47
    |    |--C. (C.) extensus Viets 1940N47
    |    |--C. (C.) gibberipes Viets 1936N47
    |    |--C. (C.) glyptoderma Trouessart 1888N47
    |    |--C. (C.) gracilis (Viets 1936) [=Copidognathopsis gracilis]N47
    |    |--C. (C.) hummelincki (Viets 1936) [=Copidognathopsis hummelincki]N47
    |    |--C. (C.) johnstoni (Womersley 1937) [=Werthella johnstoni]N47
    |    |--C. (C.) lamellosus (Lohmann 1893) [=Halacarus lamellosus]H98
    |    |--C. (C.) latus Viets 1927N47
    |    |--C. (C.) magnipalpus (Police 1909)N47
    |    |    |--C. m. magnipalpusN47
    |    |    |--C. m. ponticus Viets 1936N47
    |    |    `--C. m. serratisetus Viets 1940N47
    |    |--C. (C.) pachypus Newell 1947N47
    |    |--C. (C.) poucheti (Trouessart 1893) [=Halacarus poucheti]N47
    |    |--C. (C.) pulcher (Lohmann 1893) [=Halacarus pulcher]N47
    |    |--C. (C.) punctatissimus (Gimbel 1919) [=Halacarus (C.) punctatissimus]N47
    |    |--C. (C.) reticulatus (Trouessart 1894) [=Halacarus (C.) reticulatus]T05
    |    |--C. (C.) septentrionalis (Halbert 1915) [=Halacarus (C.) septentrionalis]N47
    |    |--C. (C.) simonisN47
    |    |--C. (C.) tricorneata Lohmann 1938N47
    |    `--C. (C.) triops Viets 1936N47
    `--C. (Arhodeoporus Newell 1947)N47
         |--C. (*A.) arenarius Newell 1947N47
         |--‘Arhodeoporus’ disparilis Bartsch 1997H98
         |--C. (A.) gracilipes (Trouessart 1889) [=C. (Copidognathopsis) gracilipes]N47
         |    |--C. g. gracilipesN47
         |    |--C. g. largiforatus (Trouessart 1899) [=C. (Copidognathopsis) gracilipes largiforatus]N47
         |    `--C. g. quadricostatus (Trouessart 1894) [=C. (Copidognathopsis) gracilipes quadricostatus]N47
         |--‘Arhodeoporus’ leptopus Bartsch 1997H98
         |--‘Arhodeoporus’ psammophilus Bartsch 1993H98
         |--C. (A.) submarinus Newell 1947N47
         `--‘Arhodeoporus’ wadjemupis Bartsch 1997H98
Copidognathus incertae sedis:
  C. alvinusWP99
  C. ampliatus Bartsch 1994H98
  C. australensis (Lohmann 1909) [=Halacarus australensis]H98
  C. bispinus Bartsch 1994H98
  C. bistriatus Bartsch 1994H98
  C. caelatus Bartsch 1994H98
  C. canaliculifer Bartsch 1994H98
  C. crassispinus Bartsch 1994H98
  C. cribellus Bartsch 1993H98
  C. curtusK09
  C. dubiosus Bartsch 1994H98
  C. fabriciusiM62
  C. gibboides Bartsch 1977B86
  ‘Copidognathopsis’ gibbusM62
  C. hartmanni Bartsch 1972B86
  C. humerosusM62
  C. laeviusculus Bartsch 1993H98
  C. laminifer Bartsch 1994H98
  C. lepidus Bartsch 1977B86
  C. lineatus Bartsch 1977B86
  C. lohmanni Trouessart 1889L90
  C. lubricus Bartsch 1986L90
  C. majorinus Bartsch 1993H98
  C. multiporus Bartsch 1994H98
  C. nasutus Bartsch 1994H98
  C. nautileiWL09
  C. papillatusWL09
  C. paucisporus Bartsch 1977B86
  C. punctellus Bartsch 1994H98
  C. pygmaeus Bartsch 1980B86
  C. rhodostigma [=Halacarus (Copidognathus) rhodostigma]T96
  C. scuna Otto 1994H98
  C. strigellus Bartsch 1994H98
  C. tabellio [=Halacarus (Copidognathus) tabellio]T96
  C. tectiporusWL09

*Type species of generic name indicated


[B86] Bartsch, I. 1986. Acari: Halacaridae. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 638–642. E. J. Brill/Dr W. Backhuys: Leiden.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[K09] Krantz, G. W. 2009. Form and function. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 5–53. Texas Tech University Press.

[L90] Luxton, M. 1990. The marine littoral mites of the New Zealand region. Journal of the Royal Society of New Zealand 20 (4): 367–418.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[N47] Newell, I. M. 1947. A systematic and ecological study of the Halacaridae of eastern North America. Bulletin of the Bingham Oceanographic Collection 10 (3): 1–232.

[T05] Trägårdh, I. 1905. Monographie der arktischen Acariden. In: Römer, F., & F. Schaudinn (eds) Fauna Arctica. Eine Zusammenstellun der arktischen Tierformen, mit besonder Berücksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nördliche Eismeer im Jahre 1898 vol. 4 pp. 1–78. Gustav Fischer: Jena.

[T96] Trouessart, E. 1896. Resultats scientifiques de la campagne du “Caudan” dans le Golfe de Gascogne: Halacariens. Annales de l’Universite de Lyon 26: 325–353, pls 9–11.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 233–420. Texas Tech University Press.

[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).

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