Neides tipularius, copyright Roger Key.

Belongs within: Panheteroptera.
Contains: Pyrrhocoroidea, Coreidae, Alydidae.

The Coreoidea are a group of plant-feeding bugs, most species of which possess ocelli and a series of longitudinal accessory veins in the forewing membrane (Carver et al. 1991). Members of this group include the Berytidae, stilt bugs, extremely slender and long-legged bugs characterised by short and spindle-shaped apical segments on the antennae. The Colobathristidae are also slender and long-legged (albeit not as remarkably so as the Berytidae) with narrow wings that have the venation on the membrane reduced or absent. The Piesmatidae are small bugs with a reduced membrane on the hemelytra, with the pronotum and corium being densely punctate. The Hyocephalidae are a group of seed-eating bugs endemic to Australia characterised by the presence of ovoid, pore-bearing plates ventrolaterally at the base of the abdomen. Idiostolidae are flattened, ground-living bugs found only in Nothofagus forests of Australia and South America (Carver et al. 1991).

Characters (from Carver et al. 1991): Macropterous or brachypterous; brachypters with antennifers inserted dorsolaterally; ocelli present; membrane of hemelytron in macropters with at least 6 and usually many more longitudinal veins; S6 with 3 or fewer trichobothria on each side.

    |  i. s.: Heterogastridae [Heterogastrinae]SE03
    |           |--DinomachusMC13
    |           |--Woodwardothignus gracilisCGW91
    |           `--Heterogaster Schilling 1829SE03
    |                `--H. urticae (Fabricius 1775)SE03
    |  |    |--DayakiellinaeSP02
    |  |    `--Phenacantha [Colobathristinae]CGW91
    |  |         |--P. australiaeCGW91
    |  |         |--P. pacifica Horváth 1904K08
    |  |         |--P. saccharicidaRD77
    |  |         `--P. viridipennisYR12
    |  `--+--Berytidae [Berytinae, Neididae]YR12
    |     |    |--Berytus clavipesH01
    |     |    |--Xenoloma Harris 1943H96
    |     |    |--Phaconotus Harris 1943H96
    |     |    |--Protacanthus Uhler 1893D01
    |     |    |    `--P. haleiCGW91
    |     |    |--Diabolonotus Henry 1996H96
    |     |    |    `--*D. pilosus Henry 1996H96
    |     |    |--Oedalocanthus Henry 1996H96
    |     |    |    `--*O. ornatus Henry 1996H96
    |     |    |--Metacanthus Costa 1848D01
    |     |    |    |--M. decorus (Uhler 1893) [=Protacanthus decorus; incl. M. capitatus Uhler 1894]D01
    |     |    |    `--M. plutoCGW91
    |     |    |--NeidesCGW91
    |     |    |    |--N. clavipes [=Gerris clavipes]L02
    |     |    |    |--N. maipongaG75
    |     |    |    |--N. propinquus Horváth 1901H01
    |     |    |    `--N. tipulariusH01 [=Gerris tipulariusL02]
    |     |    `--Gampsocoris pulchellusYR12
    |     `--Malcus [Malcidae]YR12
    |          `--M. setosusYR12
          |    |--IdiostolidaeYR12
          |    |    |--Idiostolus insularisYR12
          |    |    |--Monteithocoris hirsutusCGW91
          |    |    `--Trisecus pictusWEE70
          |    `--Pachymeridiidae [Hypocimicidae, Psychrocoridae, Sisyrocoridae]YR12
          |         |--Pachycoridium letum Popov 1986RJ93
          |         `--Beipiaocoris multifurcusYR12
             |  `--AlydidaeYR12
                |    |--Liorhyssus hyalinusCGW91
                |    |--Rhopalus latusYR12
                |    |--Chorosoma macilentumRD77, H01
                |    |--MyrmusRD77
                |    |    |--M. glabellus Horváth 1901H01
                |    |    |--M. longiventrisH01
                |    |    `--M. miriformisH01
                |    |--Leptocoris [Serinethinae]CGW91
                |    |    |--L. augurK08
                |    |    |--L. insularis Kirkaldy 1908K08
                |    |    |--L. mitellatusCGW91
                |    |    `--L. rufomarginatusCGW91
                |    `--Miracorizus punctatusYR12
                   |    |--Dicranocephalus setulosusYR12
                   |    `--StenocephalusRD77
                        |--Hyocephalus aprugnusYR12
                        `--Maevius indecorusCGW91

*Type species of generic name indicated


[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).

[D01] Distant, W. L. 1901. Rhynchotal notes.—IX. Heteroptera: fam. Coreidae. Annals and Magazine of Natural History, series 7, 7: 416–432.

[G75] Gross, G. F. 1975. Plant-feeding and Other Bugs (Hemiptera) of South Australia. Heteroptera—Part I. Handbook of the Flora and Fauna of South Australia.

[H96] Henry, T. J. 1996. Two new genera and two new species of New World stilt bugs (Heteroptera: Berytidae). Proceedings of the Entomological Society of Washington 98 (3): 533–540.

[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[MC13] Majer, J. D., S. K. Callan, K. Edwards, N. R. Gunawardene & C. K. Taylor. 2013. Baseline survey of the terrestrial invertebrate fauna of Barrow Island. Records of the Western Australian Museum Supplement 83: 13–112.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[SE03] Scudder, G. G. E., & A. C. Eyles. 2003. Heterogaster urticae (Hemiptera: Heterogastridae), a new alien species and family to New Zealand. Weta 25: 8–13.

[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815+Heteroptera Latreille, 1810). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 143–157. Kluwer Academic Publishers: Dordrecht.

[WEE70] Woodward, T. E., J. W. Evans & V. F. Eastop. 1970. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 387–457. Melbourne University Press.

[YR12] Yao, Y., D. Ren, D. A. Rider & W. Cai. 2012. Phylogeny of the infraorder Pentatomomorpha based on fossil and extant morphology, with description of a new fossil family from China. PLoS One 7 (5): e37289.

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