Corynexochida

 Reconstructed exoskeletal components of Peichiashania secunda, from Shergold (1980).

Belongs within: Trilobita.
Contains: Leiostegiidae, Pagodiidae, Illaenuridae.

The Leiostegiacea are a Middle Cambrian to Lower Ordovician group of trilobites. The position of the Missisquoiidae among trilobites has been the subject of some debate; cranidial characters suggest a relationship to Leiostegiidae, though the pygidia of the two taxa are somewhat different (Lee & Chatterton 2007).

<==CorynexochidaP98
    |  i. s.: AlbertellaP79
    |         GlossopleuraP79
    |         Bathyuriscus fimbriatusP79, NS93
    |         AmphotonP79
    |         BasidechenellaP98
    |           |--B. kayseriH79
    |           `--B. lucasensisP98
    |         FieldaspisF90
    |--Leiostegiina [Leiostegiacea]RC93
    |    |  i. s.: Palocorona Shergold 1980F90
    |    |         AnnamitellaF90
    |    |--LeiostegiidaeS80
    |    |--PagodiidaeRC93
    |    |--CheilocephalidaeRC93
    |    |--IllaenuridaeRC93
    |    |--ShirakiellidaeRC93
    |    `--MissisquoiidaeS80
    |         |--Parakoldinioida Endo in Endo & Resser 1937S80
    |         |    |--*P. typicalis Endo in Endo & Resser 1937S80
    |         |    `--P. bigranulosa Shergold 1975S80
    |         `--MissisquoiaLM05
    |              |--M. depressaCT83
    |              |--M. enigmaticaLM05
    |              |--M. inflataLM05
    |              `--M. typicalisLM05
    |--Scutelluina [Illaenacea, Illaenida, Illaenina]RC93
    |    |  i. s.: PanderiaJ79
    |    |         HarpillaenusFO99
    |    |         BumastusP98
    |    |           |--B. insignisP98
    |    |           `--B. ioxusEB01
    |    |--IllaenusF71 [IllaenidaeRC93]
    |    |    `--I. johnstoni Etheridge 1896F71
    |    |--Phillipsinella [Phillipsinellidae]RC93
    |    |    |--P. matutina Dean 1973RC93
    |    |    `--P. parabola (Barrande 1846)RC93
    |    |--Styginidae [Scutelluidae]RC93
    |    |    |--TelopeltisL-AF12
    |    |    |--ScabriscutellumRC93
    |    |    `--RaymondaspisRC93
    |    |         |--R. limbataRC93
    |    |         `--R. reticulataF90
    |    `--Tsinania [Tsinaniidae]RC93
    |         |--T. canensCT83
    |         |--T. ceresCT83
    |         |--T. humilisCT83
    |         |--T. longaCT83
    |         |--T. longicephalaCT83
    |         `--T. (Tsinania) nomasRC93
    `--CorynexochinaRC93
         |--Longduia [Longduiidae]RC93
         |--DinesidaeRC93
         |--CheiruroididaeRC93
         |    |--Hunanocephalus ovalisRC93
         |    `--Cheiruroides arcticus Chernysheva 1962RC93
         |--ChenghuiidaeRC93
         |    |--ChengkouiaRC93
         |    `--XuigiellaRC93
         |--Edelsteinaspis [Edelsteinaspidae]RC93
         |    |--E. gracilisRC93
         |    `--E. ornataRC93
         |--JakutidaeRC93
         |    |--Malykania gribovae Suvorova 1960RC93
         |    `--Jakutus amplus Egorova 1976RC93
         |--OryctocephalidaeRC93
         |    |--Arthricocephalus chauveauiRC93
         |    `--Tonkinella kobayashiRC93
         |--Ogygopsis [Ogygopsidae]RC93
         |    |--O. batis (Walcott 1889)RC93
         |    `--O. klotziWB85
         |--DorypygidaeRC93
         |    |--KooteniaRC93
         |    |--DorypygeRC93
         |    `--OlenoidesRC93
         |         |--O. serratus (Rominger 1887)CB04
         |         `--O. tranans Öpik 1967RC93
         |--DolichometopidaeRC93
         |    |--HoffetellaRC93
         |    |--HemirhodonRC93
         |    `--DolichometopusT10
         |         |--D. howelliT10
         |         `--D. tateiT10
         |--ZacanthoididaeRC93
         |    |--ZacanthoidesP79
         |    |--ZacanthopsisRC93
         |    |--ZacanthopsinaRC93
         |    |--StephenaspisRC93
         |    `--ChuchiaspisRC93
         `--Corynexochidae [Corynexochacea]RC93
              |--BomnaspisRC93
              `--Corynexochus Angelin 1854SS84
                   |--*C. spinulosus Angelin 1854SS84
                   |--C. delagei Miquel 1905SS84
                   |--C. plumulaP79
                   `--C. sardous Rasetti 1972SS84

*Type species of generic name indicated

References

[CT83] Chen J.-Y. & C. Teichert. 1983. Cambrian Cephalopoda of China. Palaeontographica Abteilung A: Paläozoologie—Stratigraphie 181 (1–3): 1–102.

[CB04] Cotton, T. J., & S. J. Braddy. 2004. The phylogeny of arachnomorph arthropods and the origin of the Chelicerata. Transactions of the Royal Society of Edinburgh: Earth Sciences 94: 169-193.

[EB01] Eckert, J. D., & C. E. Brett. 2001. Early Silurian (Llandovery) crinoids from the Lower Clinton Group, western New York State. Bulletins of American Paleontology 360: 1–88.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[F90] Fortey, R. A. 1990. Ontogeny, hypostome attachment and trilobite classification. Palaeontology 33 (3): 529–576.

[FO99] Fortey, R. A. & R. M. Owens. 1999. Feeding habits in trilobites. Palaeontology 42 (3): 429–465.

[H79] House, M. R. 1979. Devonian in the Eastern Hemisphere. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A183–A217. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[J79] Jaanusson, V. 1979. Ordovician. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A136–A166. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[LM05] Lehnert, O., J. F. Miller, S. A. Leslie, J. E. Repetski & R. L. Ethington. 2005. Cambro-Ordovician sea-level fluctuations and sequence boundaries: the missing record and the evolution of new taxa. Special Papers in Palaeontology 73: 117–134.

[L-AF12] Lerosey-Aubril, R., & R. Feist. 2012. Quantitative approach to diversity and decline in late Palaeozoic trilobites. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 535–555. Springer.

[NS93] Nudds, J. R., & J. J. Sepkoski Jr. 1993. Coelenterata. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 101–124. Chapman & Hall: London.

[P79] Palmer, A. R. 1979. Cambrian. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A119–A135. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[P98] Prothero, D. R. 1998. Bringing Fossils to Life: An introduction to paleobiology. WCB McGraw-Hill: Boston.

[RC93] Romano, M., W. T. Chang, W. T. Dean, G. D. Edgecombe, R. A. Fortey, D. J. Holloway, P. D. Lane, A. W. Owen, R. M. Owens, A. R. Palmer, A. W. A. Rushton, J. H. Shergold, D. J. Siveter & M. A. Whyte. 1993. Arthropoda (Trilobita). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 279–296. Chapman & Hall: London.

[S80] Shergold, J. H. 1980. Late Cambrian trilobites from the Chatsworth Limestone, western Queensland. Bureau of Mineral Resources, Geology and GeophysicsBulletin 186: 1–111.

[SS84] Shergold, J. H., & K. Sdzuy. 1984. Cambrian and early Tremadocian trilobites from Sultan Dağ, central Turkey. Senckenbergiana Lethaea 65 (1–3): 51–135.

[T10] Taylor, T. G. 1910. The Archaeocyathinae from the Cambrian of South Australia with an account of the morphology and affinities of the whole class. Memoirs of the Royal Society of South Australia 2 (2): 55–188.

[WB85] Whittington, H. B., & D. E. G. Briggs. 1985. The largest Cambrian animal, Anomalocaris, Burgess Shale, British Columbia. Philosophical Transactions of the Royal Society of London 309: 569–609.

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