Crangonyctoidea

 Pseudocrangonyx asiaticus, from Holsinger (1986).

Belongs within: Senticaudata.
Contains: Niphargus, Neoniphargidae, Paramelitidae, Crangonyx, Stygobromus, Synurella.

The Crangonyctoidea are a group of freshwater amphipods in which the first urosomite bears a distoventral robust seta (Lowry & Myers 2013). Many families in the group contain species living in hypogean (underground, i.e. caves or groundwater) habitats and some are entirely hypogean. The majority inhabit Gondwanan landmasses with the exception of the Holarctic Crangonyctidae and the Palaearctic Niphargidae and Pseudocrangonyctidae (Lowry & Myers 2013).

Crangonyctoidea [Niphargoidea]
    |--+--+--Sandro Karaman & Barnard 1979 [Sandroidae]LM13
    |  |  |    `--*S. starmuehlneri (Ruffo 1960)B83
    |  |  `--AustroniphargidaeLM13
    |  |       |--Libertinia Iannilli, Krapp & Ruffo 2011LM13
    |  |       |--Davidia Iannilli, Krapp & Ruffo 2011LM13
    |  |       `--Austroniphargus Monod 1925LM13
    |  |            |--*A. bryophilus (Monod 1925)B83 [=Niphargopsis bryophilusB77]
    |  |            |--A. petitiW86
    |  |            `--A. starmuehlneri Ruffo 1960B77
    |  `--NiphargidaeLM13
    |       |--NiphargusLM13
    |       |--Foroniphargus Karaman 1985LM13
    |       |--Niphargopsis Chevreux 1922LM13
    |       |    `--*N. casparyi (Pratz 1866) [=Niphargus casparyi]B77
    |       |--Microniphargus Schellenberg 1934LM13
    |       |    `--*M. leruthi Schellenberg 1934B77
    |       |--Pontoniphargus Dancau 1970LM13
    |       |    `--*P. racovitzai Dancau 1970B77
    |       |--Carinurella Sket 1971LM13
    |       |    `--*C. paradoxa (Sket 1964) [=Niphargus paradoxa]B77
    |       |--Niphargobates Sket 1981LM13
    |       |    `--N. orophobata Sket 1981KR86
    |       |--Haploginglymus Mateus & Mateus 1958LM13
    |       |    |--H. bragai Mateus & Mateus 1958KR86
    |       |    `--H. lobatus Stock 1980KR86
    |       `--Niphargellus Schellenberg 1938LM13
    |            |--*N. arndti (Schellenberg 1933)B77, KR86 [=Niphargus arndtiB77]
    |            |--N. glenniei (Spooner 1952)KR86
    |            `--N. nolli Schellenberg 1938KR86
    `--+--+--+--Uronyctus Stock & Iliffe 1990 [Uronyctidae]LM13
       |  |  `--Giniphargus Williams & Barnard 1988 [Giniphargidae]LM13
       |  |       `--*G. pulchellus (Sayce 1900)B83, W86 [=Niphargus pulchellusW86]
       |  `--+--Kotumsaria Messouli, Holsinger & Reddy 2007 [Kotumsaridae]LM13
       |     `--Paracrangonyx Stebbing 1899 [Paracrangonyctidae]LM13
       |          `--*P. compactus (Chilton 1882)B83 [=Crangonyx compactusB77]
       `--+--Chillagoe Barnard & Williams 1995 [Chillagoeidae]LM13
          `--+--+--NeoniphargidaeLM13
             |  `--+--ParamelitidaeLM13
             |     `--Perthia Straškraba 1964 [Perthiidae]LM13
             |          |--*P. branchialis (Nicholls 1924) [=Neoniphargus branchialis]B77
             |          `--P. acutitelsonTJ06
             `--+--Sternophysinx Holsinger & Straškraba 1973 [Sternophysingidae]LM13
                |    |--*S. robertsi (Methuen 1911)B83, W86 [=Eucrangonyx robertsiB77]
                |    |--S. alca Griffiths 1981W86
                |    `--S. filaris Holsinger & Straskraba 1973W86
                `--+--PseudocrangonyctidaeLM13
                   |    |--Procrangonyx Schellenberg 1934LM13 [=Eocrangonyx Schellenberg 1937B83]
                   |    |    `--*P. japonicus (Ueno 1930) [=Eucrangonyx japonicus, *Eocrangonyx japonicus]H86
                   |    `--Pseudocrangonyx Akatsuka & Komai 1922LM13 [incl. Niphargonyx Derzhavin 1927H86]
                   |         |--*P. yezonis Akatsuko & Komai 1922B83
                   |         |--P. asiaticus Ueno 1934H86
                   |         |--P. bohaensis (Derzhavin 1927) [=*Niphargonyx bohaensis]H86
                   |         |--P. camtschaticus Birstein 1955H86
                   |         |--P. coreanus Ueno 1966H86
                   |         |--P. kyotonis Akatsuka & Komai 1922H86
                   |         |--P. levanidovi Birstein 1955H86
                   |         |--P. manchuricus Oguro 1938H86
                   |         `--P. shikokunis Akatsuka & Komai 1922H86
                   `--CrangonyctidaeLM13
                        |--CrangonyxLM13
                        |--StygobromusLM13
                        |--SynurellaLM13
                        |--Pachypodacrangonyx Boutin & Coineau 1987LM13
                        |--Stygonyx Bousfield & Holsinger 1989LM13
                        |--Lyurella Derzhavin 1939LM13
                        |    `--*L. hyrcana Derzhavin 1939B77
                        |--Stygonectes Hay 1902B77
                        |    `--*S. flagellatus Benedict 1896B77
                        |--Bactrurus Hay 1902LM13
                        |    |--*B. mucronatus (Forbes 1876)B77, H86 [=Eucrangonyx mucronatusB77]
                        |    |--B. brachycaudus Hubricht & Mackin 1940H86
                        |    `--B. hubrichti Shoemaker 1945H86
                        `--Palaeogammarus Zaddach 1864H86, B77
                             |--*P. sambiensis Zaddach 1864B77
                             |--P. balticus Lucks 1928BWW93
                             |--P. danicus Just 1974BWW93
                             `--P. sambiensis Zaddach 1864BWW93

*Type species of generic name indicated

References

[B77] Bousfield, E. L. 1977. A new look at the systematics of gammaroidean amphipods of the world. Crustaceana Suppl. 4: 282–316.

[B83] Bousfield, E. L. 1983. An updated phyletic classification and palaeohistory of the Amphipoda. In: Schram, F. R. (ed.) Crustacean Phylogeny pp. 257–277. A. A. Balkema.

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[H86] Holsinger, J. R. 1986. Holarctic crangonyctid amphipods. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 535–549. E. J. Brill/Dr W. Backhuys: Leiden.

[KR86] Karaman, G. S., & S. Ruffo. 1986. Amphipoda: Niphargus-group (Niphargidae sensu Bousfield, 1982). In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 514–534. E. J. Brill/Dr W. Backhuys: Leiden.

[LM13] Lowry, J. K., & A. A. Myers. 2013. A phylogeny and classification of the Senticaudata subord. nov. (Crustacea: Amphipoda). Zootaxa 3610: 1–80.

[W86] Williams, W. D. 1986. Amphipoda on land-masses derived from Gondwana. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 553–559. E. J. Brill/Dr W. Backhuys: Leiden.

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