Nagurus cristatus, copyright Arp Kruihof.

Belongs within: Oniscidea.
Contains: Armadillidae, Porcellio, Oniscidae, Philosciidae, .

The Crinocheta are a group of terrestrial isopods characterised by an ’embolophore’ type of male genitalia with the two vasa deferentia opening by separate pores on either side of the penis towards its apex (Hatch 1947, as Embolophora). They are divided between the Armadilloidea, in which the pleopods carry respiratory pseudotracheae, and the Oniscoidea in which such structures are absent.

<==Crinocheta [Embolophora]MD01
    |    |--AtlantididaeMD01
    |    |--BalloniscidaeMD01
    |    |--PeriscyphicidaeMD01
    |    |--ArmadillidaeMD01
    |    |--EubelidaeMD01
    |    |    |--Eubelum rusingaense Morris 1979BWW93
    |    |    `--ElumoidesJG19
    |    |--CylisticusM90 [CylisticidaeMD01]
    |    |    |--C. convexusM90 [incl. C. convexus var. confluentusH47, Porcellio glaberH47]
    |    |    `--C. rotabilisD01
    |    |--Actaecia Dana 1853V73 [incl. Cylloma Budde-Lund 1885V73; ActaeciidaeMD01]
    |    |    |--A. euchroa Chilton 1901NB26
    |    |    |--A. opihensis Chilton 1901NB26
    |    |    `--A. pallida Nicholls & Barnes 1926V73
    |    |--ArmadillidiidaeMD01
    |    |    |--Protosphaeoniscus tertiarius Schmalfuss 1980BWW93
    |    |    `--ArmadillidiumM90
    |    |         |--A. (Armadillidium) vulgare [incl. Armadillo pilularis Say 1818]H47
    |    |         `--A. (Pseudosphaerium) nasatumH47
    |    |--PorcellionidaeMD01
    |    |    |--PorcellioS77
    |    |    |--Agabiformius lentus (Budde-Lund 1885)V73
    |    |    `--Porcellionides Miers 1877JP13
    |    |         `--P. pruinosus (Brandt 1833)JP13 [=Metoponorthus pruinosusM90; incl. P. immaculatusH47]
    |    `--Trachelopodidae [Trachelipidae]MD01
    |         |--Trachelipus [incl. Tracheoniscus]M90
    |         |    `--T. rathkeiM90 (see below for synonymy)
    |         `--Nagurus Holthuis 1949 [incl. Nagara Budde-Lund 1908 (preoc.), Nagaroides Wahrberg 1922]V73
    |              |  i. s.: N. aegaeus Schmalfuss 1977S77
    |              |         N. carinatus [incl. N. hermonensis Vandel 1955]S77
    |              |         N. declivus (Wahrberg 1922)V73
    |              |         N. gracillimus (Wahrberg 1922)V73
    |              |         N. longiflagellatus (Wahrberg 1922)V73
    |              |         ‘Nagara’ modestaWKN94
    |              |         N. rhodiensisS77
    |              |         N. tumidus (Wahrberg 1922)V73
    |              `--N. (Nagurus)H83
    |                   |--N. (N.) cristatus (Dollfus 1889)H83, V73
    |                   `--N. (N.) nanusH83
         |  i. s.: Tasmanoniscus Vandel 1973V73
         |           `--*T. evansi Vandel 1973V73
         |--Olibrinus Budde-Lund 1913JP13 [OlibrinidaeMD01]
         |--ProtosphaeroniscusP92 [SphaeroniscidaeMD01]
         |    `--P. tertiariusP92
         |--HalophilosciaPP64 [HalophilosciidaeMD01]
         |    `--H. couchiiPP64
         |--Australoniscus Vandel 1973V73 [BathytropidaeMD01]
         |    `--*A. springetti Vandel 1973V73
         |--DetoV73 [DetonidaeMD01]
         |    `--D. marina (Chilton 1884)V73
         |--RhyscotusM90 [RhyscotidaeMD01]
         |    `--R. texensisM90
         |    |--PlatyarthrusM90
         |    |    |--P. hoffmannseggiM90
         |    |    `--P. schoebliM90
         |    `--Trichorhina [Squamiferidae]P92
         |         |--T. australiensis Wahrberg 1922V73
         |         |--T. heterophthalmaM90
         |         |--T. tomentosaM90
         |         `--T. triocisM90
              |--Scyphacella arenicolaM90
              |--Detonella [Scyphacinae]H47
              |    |--D. papillicornis [=Trichoniscus papillicornis; incl. D. lohmanderi]H47
              |    `--D. sachalinaH47
              |    |--H. searli Chilton 1920 [incl. Philoscia salina Baker 1926]V73
              |    `--H. stephensi Nicholls & Barnes 1927V73
              `--Armadilloniscus [Armadilloniscinae]H47
                   |--A. ellipticusM90
                   |--A. holmesiM90
                   |--A. lindahliM90
                   `--A. tuberculatus [=Actoniscus tuberculatus]H47
Nomen nudum: Australoniscus alticolus Vandel 1973V73

Trachelipus rathkeiM90 [incl. Porcellio dorsalisH47, P. lateralisH47, P. limatusH47, P. limbalisH47, P. marginatusH47, P. mixtusH47, P. multiguttatusH47, T. rathkei var. ochraceusH47, Porcellio variegatusH47, T. rathkei var. variusH47]

*Type species of generic name indicated


[BW04] Balke, M., C. H. S. Watts, S. J. B. Cooper, W. F. Humphreys & A. P. Vogler. 2004. A highly modified stygobiont diving beetle of the genus Copelatus (Coleoptera, Dytiscidae): taxonomy and cladistic analysis based on mitochondrial DNA sequences. Systematic Entomology 29: 59–67.

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[D01] Dollfus, A. 1901. Isopodák [Isopodes]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 371–374. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[H83] Hammen, L. van der. 1983. Contribution to the knowledge of the soil-fauna of New Guinea. Zoologische Verhandelingen 206: 1–36.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.

[JP13] Judd, S., & G. Perina. 2013. An illustrated key to the morphospecies of terrestrial isopods (Crustacea: Oniscidea) of Barrow Island, Western Australia. Records of the Western Australian Museum Supplement 83: 185–207.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[M90] Muchmore, W. B. 1990. Terrestrial Isopoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 805–817. John Wiley & Sones: New York.

[NB26] Nicholls, G. E., & H. M. Barnes. 1926. A description of two new terrestrial isopods from Western Australia. Journal of the Royal Society of Western Australia 12 (17): 149–159.

[PP64] Peres, J. M., & J. Picard. 1964. Nouveau manuel de bionomie benthique de la mer Mediterranee. Recueil des Travaux de la Station Marine d’Endoume, Bulletin 31 (27): 5–137.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[S77] Schmalfuss, H. 1977. Eine neue Nagurus-Art aus der Süd-Agäis (Crustacea: Isopoda: Oniscoidea: Trachelipidae). Senckenbergiana Biologica 57 (4–6): 359–365.

[V73] Vandel, A. 1973. Les isopodes terrestres de l’Australie. Étude systématique et biogéographique. Mémoires du Muséum National d’Histoire Naturelle, Série A, Zoologie 82: 1–171.

[WKN94] Wrensch, D. L., J. B. Kethley & R. A. Norton. 1994. Cytogenetics of holokinetic chromosomes and inverted meiosis: keys to the evolutionary success of mites, with generalizations on eukaryotes. In: M. A. Houck (ed.) Mites: Ecological and Evolutionary Analyses of Life-history Patterns pp. 282–343. Chapman & Hall: New York.

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